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harlequin lotus, seaside bird's-foot lotus, seaside bird's-foot-trefoil, seaside lotus, witches teeth

big deervetch, broad leaf lotus

Habit Herbs usually low, 10–40 cm, glabrous; stoloniferous or rhizomatous. Herbs 70–150 cm, villous or sparsely strigulose to glabrate; root-spreading.
Stems

1–20+, often decumbent, sometimes ascending, unbranched, fleshy, base often spongy.

(1–)5–20+, ascending to erect, fleshy, stout, hollow.

Leaves

stipules lanceolate-ovate to ovate or deltate, 2–8 mm, ± scarious, fragile; petiolate to long-petiolate;

rachis ± straight, 1–4 cm;

leaflets (3–)5–7, petiolulate, blades elliptic to obovate, 6–20 mm, apex obtuse to emarginate, surfaces glabrous.

stipules triangular or ovate to lanceolate, (2–)5–8 mm, membranous becoming scarious, often becoming inconspicuous; short-petiolate to petiolate;

rachis often curved, 6–12 cm;

leaflets (7–)9–15(–19), petiolulate, blades widely ovate to widely elliptic or widely obovate, (10–)20–30 mm, apex obtuse to emarginate, surfaces sparsely strigillose to glabrate.

Peduncles

ascending to spreading, 2.3–9 cm, equaling or longer than subtending leaf, elongating in fruit, glabrous;

bract subtending umbel, 1–3(–7)-foliolate.

ascending, 3–8 cm, usually shorter than, rarely equaling, subtending leaf, elongating in fruit, glabrate or glabrous;

bract medial or absent, leaflets (1 or)3 or 5-foliolate.

Umbels

(2 or)3–9-flowered.

8–20-flowered.

Flowers

10–16 mm;

calyx reddish, 4.5–6 mm, glabrous, tube cylindric, 2.5–4 mm, lobes subulate to narrowly triangular, 2–2.5 mm, eciliate;

corolla banner yellow, wings and keel pale to dark pink, sometimes whitish, wings fading white, keel pink- to purple-tipped, wings ± equaling or longer than keel, claw longer than calyx tube, banner reflexed to 180°, keel apex acute to subacute.

asymmetric, 12–16(–18) mm;

calyx green marked with purplish or pinkish, or purplish to pinkish, 4.5–8 mm, glabrous, tube cylindric, 4–6 mm, lobes deltate, 0.5–2 mm, sparsely ciliolate or eciliate;

corolla greenish or whitish becoming marked with red or purple, wings ± equaling keel, claw ± longer than calyx tube, banner implicate-ascending to 45°, keel apex acute.

Legumes

± ascending to ± inclined, brown, oblong, turgid, 20–30 × 2–3 mm, thinly leathery, incompletely septate, glabrous.

± ascending to inclined, reddish to brown, linear, turgid, (10–)35–70 × 3–5 mm, leathery, incompletely septate, glabrous.

Seeds

(8–)10–14(–16), olive brown to brown, oblong, 1–1.5 mm.

7–12, olive or reddish to dark brown, obovoid, 3–4 mm.

2n

= 14.

= 14.

Hosackia gracilis

Hosackia crassifolia

Phenology Flowering spring(–early summer).
Habitat Water or springy areas, shores, coastal meadows, wet meadows or woodlands, sphagnum bogs, pastures, roadside ditches, coastal areas.
Elevation 0–200(–700) m. (0–700(–2300) ft.)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from USDA
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Hosackia gracilis reaches its northern limit in southeastern Vancouver Island and adjacent Gulf islands near Victoria, British Columbia (G. W. Douglas and M. Ryan 2006). Southward it follows the coast of Washington (Grays Harbor County), Oregon (with an inland incursion to Lane County), and California (to San Luis Obispo County), with disjunct populations inland in the foothills of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Lotus crassifolius (Bentham) Greene [not Persoon] is an illegitimate name that pertains to Hosackia crassifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs glabrate or sparsely strigulose; ovules.
var. crassifolia
1. Herbs villous; ovules 14–21.
var. otayensis
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Hosackia Fabaceae > subfam. Faboideae > Hosackia
Sibling taxa
H. alamosana, H. crassifolia, H. incana, H. oblongifolia, H. pinnata, H. rosea, H. stipularis, H. yollabolliensis
H. alamosana, H. gracilis, H. incana, H. oblongifolia, H. pinnata, H. rosea, H. stipularis, H. yollabolliensis
Subordinate taxa
H. crassifolia var. crassifolia, H. crassifolia var. otayensis
Synonyms Anisolotus formosissimus, Lotus formosissimus
Name authority Bentham: Trans. Linn. Soc. London 17: 365. (1836) Bentham: Trans. Linn. Soc. London 17: 365. (1836)
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