Horkelia truncata
Horkelia rydbergii
Ramona horkelia
Rydberg's horkelia
erect, 2–6(–8) dm, hairs spreading.
ascending to erect, (1–)2–7 dm, hairs ascending to appressed.
planar, 4–13 × 2–3.5(–4) cm;
stipules entire or basally lobed;
leaflets 1–3 per side, separate, oblong to obovate, 10–30 × 5–12(–15) mm, ± 1/2 as wide as long, divided 1/6–1/4 to midrib into (15–)20–30 acute to obtuse teeth, sparsely short-pilose to glabrate, especially adaxially.
± planar, (4–)8–20(–30) × 0.8–2(–3) cm;
stipules entire;
leaflets 7–14 per side, separate to ± overlapping at least distally, cuneate to flabellate, (3–)5–15(–22) × 3–10(–15) mm, ± 1/2 to as wide as long, divided ± 1/3 to midrib into (3–)5–10 acute to obtuse teeth, densely (to sparsely) strigose or pilose.
4–7.
(2 or)3–6(–8).
open, flowers arranged individually.
± open to congested, flowers arranged individually or in glomerules.
4–20(–30) mm.
(1–)2–8 mm.
12–17 mm diam.;
epicalyx bractlets broadly ovate, 3–4.5 × 2–3 mm, ± 2/3 length of sepals, entire;
hypanthium 1.5–2 × 5–8 mm, less than 1/2 as deep as wide, interior glabrous;
sepals ± reflexed, broadly ovate, 3.5–5.5 mm;
petals obovate to round, 5–7 × (3–)4–7 mm, apex ± rounded;
filaments 1–2 × 0.5–2 mm, anthers 0.8–1 mm;
carpels 50–80;
styles 1.5–3 mm.
10–15 mm diam.;
epicalyx bractlets lanceolate to narrowly elliptic, 2–3.5(–4) × 0.5–1 mm, ± 3/4 length of sepals, entire;
hypanthium 1–1.5(–2) × 2.5–4.5(–6) mm, less than 1/2 as deep as wide, interior pilose;
sepals spreading to ± reflexed, lanceolate, (2.5–)3–5(–5.2) mm;
petals oblong to oblanceolate, 4–5.5 × 1.2–2.5 mm, apex rounded to truncate or slightly emarginate;
filaments 0.5–2 × 0.5–1 mm, anthers 0.6–1 mm;
carpels 20–50(–120);
styles 2–4 mm.
brown, 1.3–1.5 mm, smooth.
light to dark brown, 1–1.5 mm, smooth or roughened.
Horkelia truncata
Horkelia rydbergii
Of conservation concern.
Horkelia truncata is confined to the Peninsular Ranges of San Diego County, California, and northern Baja California, Mexico. Plants of the species are unusual in Horkelia in that the terminal leaflet is usually notably larger than the lateral leaflets, unlobed, and distinct, rather than confluent with the distal lateral ones.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Horkelia rydbergii occurs in the Transverse Ranges of Kern, Los Angeles, San Bernardino, and Ventura counties. Reports from farther north (as H. bolanderi var. parryi) are referable to H. marinensis, H. yadonii, or, possibly, H. cuneata. Although there has been occasional nomenclatural confusion between this taxon (as H. bolanderi var. parryi) and H. parryi Greene, the two are taxonomically and geographically distinct.
Plants in the Mount Pinos–Lockwood Valley area of Kern and Ventura counties, and on Frazier Mountain, are distinctly gray with an abundance of tightly curled hairs. The leaves tend to be relatively short (to 8 cm) with relatively small leaflets (3–6 mm) on the basal leaves. Stems are relatively short (to 3 dm), and the fruits are dark brown, distinctly rugose, and 1.3–1.5 mm. This phase (including the type of Horkelia rydbergii) abruptly gives way to a grayish green phase with a slightly less dense, pilose indument in the San Bernardino and San Gabriel mountains. Here the plants tend to have larger basal leaves (to 30 cm) and leaflets (5–12 mm), longer stems (to 7 dm), and fruits that are a lighter brown, mostly smooth, and 1–1.2 mm. This phase in turn grades into even less densely hairy plants near Bear Lake, where the type of H. bolanderi var. parryi was obtained.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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