Horkelia tridentata var. flavescens |
Horkelia tridentata |
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hairycup horkelia, pale horkelia, three tooth horkelia |
cluster horkelia, three-tooth horkelia |
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Habit | Plants mostly rosette-forming, grayish. | |||||
Stems | ± decumbent, 0.5–2.5(–4.5) dm. |
decumbent to nearly erect, 0.5–4(–7) dm, hairs 1–3 mm proximally, glands absent or sparse distally. |
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Basal leaves | planar, 3–12 × 1–3(–4) cm, usually sericeous to villous, sometimes glabrate adaxially; stipules entire or rarely forked or pectinate; leaflets 2–5(or 6) per side, separate to slightly overlapping, elliptic to cuneate-oblong, (3–)5–20(–30) × 1.5–7(–10) mm, (1/4–)1/3–1/2 as wide as long, divided 1/10–1/6 or less to midrib into (0–)3(or 4) triangular teeth 1 mm wide, these restricted to apex. |
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Cauline leaves | 1–5; stipules 3–12 mm, entire or deeply 3–5-lobed, lanceolate to ovate. |
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Inflorescences | composed of usually 1 3–25-flowered ± corymbiform cluster. |
open to congested, flowers arranged in ± capitate glomerules or in ± corymbiform clusters. |
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Pedicels | (1–)2–6(–10) mm. |
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Flowers | 6–10 mm diam.; hypanthium 2.5–5 mm diam., interior pilose (except in some North Coast Ranges populations); petals ± broadly oblanceolate, (1–)1.5–3(–4) × 0.5–1.5 mm, often shorter than sepals, apex obtuse to rounded or mucronate; filaments 1–2 × 0.2–0.4(–0.8) mm, anthers 0.3–0.5 mm; styles 1.5–2.5 mm. |
4–10 mm diam.; epicalyx bractlets linear to narrowly elliptic, 0.8–2(–3) × 0.2–0.4 mm, 1/2–2/3 length of sepals; hypanthium 1–2.5 × 2–5 mm, 1/2 to nearly as deep as wide, interior glabrous or pilose; sepals moderately reflexed, abaxially green to reddish, 1.5–3 mm; petals white, sometimes tinged with pink, linear to broadly oblanceolate, (1–)1.5–4(–4.5) × 0.3–1.5 mm, apex acute to rounded or mucronate; filaments 0.5–2 × 0.1–0.4(–0.8) mm, anthers 0.2–0.5 mm; carpels 5–15; styles 1–2.5 mm. |
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Achenes | 2–2.5 mm, strongly rugose. |
light brown, 1.5–2.5 mm, rugose. |
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Horkelia tridentata var. flavescens |
Horkelia tridentata |
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Phenology | Flowering summer. | |||||
Habitat | Open areas, primarily in conifer woodlands, often on serpentine soil | |||||
Elevation | 700–2000 m (2300–6600 ft) | |||||
Distribution |
CA; OR |
CA; OR
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Discussion | Variety flavescens occurs primarily in the mountains of northwestern California, from Mendocino to Siskiyou counties. A single collection is confirmed from Curry County, Oregon (Bear Camp Trail, Leach 4554, OSC); other collections from Curry and Josephine counties may be hybrids between this variety and Horkelia congesta var. nemorosa. A second cluster of populations occurs in central Plumas and adjacent Lassen counties, California, and historic collections are known from near Truckee on the border of Nevada and Placer counties, California. D. D. Keck (1938) suggested that this bicentric distribution may have resulted from the independent coalescence of the diagnostic characteristics of var. flavescens. The locality of a collection purportedly from Washoe County, Nevada (80 miles north of Reno, Brooks s.n., RENO) needs to be confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Rydberg established Horkelia flavescens on the basis of its more congested inflorescence, purportedly yellowish petals, and broader filaments. As noted by D. D. Keck (1938), the petals are usually white, at least when fresh, and the other characters are highly variable; he reduced H. flavescens to a subspecies of H. tridentata, which is here treated at the rank of variety. In general, var. flavescens tends to be more decumbent with a single, loosely congested flower cluster and broader petals; these characters completely overlap and can vary independently. The presence or absence of hairs on the hypanthium interior is the single most reliably diagnostic character and is the primary determiner of the geographic distributions presented here. The exception is a cluster of populations from the extreme southern end of the coastal range of var. flavescens in Colusa, Lake, and Mendocino counties, California, which have glabrous hypanthia but otherwise have all the diagnostic characters of var. flavescens. Although provisionally included here within var. flavescens, these populations may be better treated as a separate variety. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 270. | FNA vol. 9, p. 269. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | H. flavescens, H. tridentata subsp. flavescens, Potentilla tilingii var. flavescens | |||||
Name authority | (Rydberg) Ertter & Reveal: Novon 17: 320. (2007) | Torrey: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 84, plate 6. (1857) | ||||
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