Horkelia tenuiloba |
Rosaceae subfam. rosoideae |
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Santa Rosa, Santa Rosa horkelia, Sonoma horkelia, thin-lobed, thin-lobed horkelia |
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Habit | Plants loosely matted, green. | Herbs, shrubs, or subshrubs. |
Stems | ascending to erect, 1–4 dm, hairs ± spreading. |
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Leaves | alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound; stipules present, rarely absent. |
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Basal leaves | weakly planar to ± cylindric, 5–15(–20) × 0.5–1.5 cm; stipules entire; leaflets 8–16(–20) per side, ± overlapping especially distally, cuneate to flabellate, 3–10 × 2–10 mm, 1/2 to nearly as wide as long, divided 1/2–3/4+ to midrib into 3–8 linear to narrowly oblanceolate or narrowly elliptic lobes, sparsely villous at least marginally, often with a tuft of hairs apically. |
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Cauline leaves | 2–5. |
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Inflorescences | open to congested, flowers arranged individually and in glomerules, these sometimes subcapitate. |
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Pedicels | 1–6 mm. |
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Flowers | 10 mm diam.; epicalyx bractlets narrowly to broadly lanceolate, 1–3 × 0.5–1 mm, slightly shorter than sepals, entire; hypanthium 1–1.2 × 2.5–4.5 mm, less than 1/2 as deep as wide, interior pilose; sepals spreading to reflexed, lanceolate, 3–5 mm; petals oblanceolate, 2.5–4.5 × 1.5 mm, apex emarginate; filaments (1–)1.5–2 × 0.5 mm, anthers 0.4–0.6 mm; carpels 10–25; styles 1.8–2.2 mm. |
torus usually enlarged, sometimes small or absent; carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae); ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula). |
Fruits | achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets; styles persistent or deciduous, not elongate (elongate but not plumose in Geum). |
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Achenes | light brown, 1.5 mm, smooth or slightly rugose. |
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x | = 7(8). |
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2n | = 28. |
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Horkelia tenuiloba |
Rosaceae subfam. rosoideae |
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Phenology | Flowering summer. | |
Habitat | Sandy soil, openings, in chaparral, oak woodlands | |
Elevation | 50–500 m (200–1600 ft) | |
Distribution |
CA
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North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia |
Discussion | Of conservation concern. Horkelia tenuiloba occurs on the western edges of the northern Coast Ranges in Marin, Mendocino, and Sonoma counties. Populations from San Luis Obispo formerly included in this species now are part of H. yadonii. A specimen (M. K. C[urran], July 5, 1885, UC) unequivocally of H. tenuiloba purportedly from San Luis Obispo is in all likelihood mislabeled with respect to locality. Horkelia tenuiloba is commonly associated with seral openings in chaparral and woodlands and might be dependent on periodic disturbance by fire. W. L. Jepson (1909–1943, vol. 2) used Potentilla stenoloba (1895) for the species encompassing the types of Horkelia tenuiloba and P. micheneri. The epithet micheneri (1893) has priority at species rank within Potentilla, since P. tenuiloba (Torrey) Greene is a later homonym of P. tenuiloba Jordan. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily. Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 252. | FNA vol. 9, p. 23. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | H. fusca var. tenuiloba, Potentilla micheneri, P. stenoloba | |
Name authority | (Torrey) A. Gray: Proc. Amer. Acad. Arts 6: 529. (1865) | Arnott: Botany, 107. (1832) |
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