Horkelia tenuiloba |
Horkelia tridentata |
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Santa Rosa, Santa Rosa horkelia, Sonoma horkelia, thin-lobed, thin-lobed horkelia |
cluster horkelia, three-tooth horkelia |
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Habit | Plants loosely matted, green. | Plants mostly rosette-forming, grayish. | ||||
Stems | ascending to erect, 1–4 dm, hairs ± spreading. |
decumbent to nearly erect, 0.5–4(–7) dm, hairs 1–3 mm proximally, glands absent or sparse distally. |
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Basal leaves | weakly planar to ± cylindric, 5–15(–20) × 0.5–1.5 cm; stipules entire; leaflets 8–16(–20) per side, ± overlapping especially distally, cuneate to flabellate, 3–10 × 2–10 mm, 1/2 to nearly as wide as long, divided 1/2–3/4+ to midrib into 3–8 linear to narrowly oblanceolate or narrowly elliptic lobes, sparsely villous at least marginally, often with a tuft of hairs apically. |
planar, 3–12 × 1–3(–4) cm, usually sericeous to villous, sometimes glabrate adaxially; stipules entire or rarely forked or pectinate; leaflets 2–5(or 6) per side, separate to slightly overlapping, elliptic to cuneate-oblong, (3–)5–20(–30) × 1.5–7(–10) mm, (1/4–)1/3–1/2 as wide as long, divided 1/10–1/6 or less to midrib into (0–)3(or 4) triangular teeth 1 mm wide, these restricted to apex. |
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Cauline leaves | 2–5. |
1–5; stipules 3–12 mm, entire or deeply 3–5-lobed, lanceolate to ovate. |
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Inflorescences | open to congested, flowers arranged individually and in glomerules, these sometimes subcapitate. |
open to congested, flowers arranged in ± capitate glomerules or in ± corymbiform clusters. |
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Pedicels | 1–6 mm. |
(1–)2–6(–10) mm. |
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Flowers | 10 mm diam.; epicalyx bractlets narrowly to broadly lanceolate, 1–3 × 0.5–1 mm, slightly shorter than sepals, entire; hypanthium 1–1.2 × 2.5–4.5 mm, less than 1/2 as deep as wide, interior pilose; sepals spreading to reflexed, lanceolate, 3–5 mm; petals oblanceolate, 2.5–4.5 × 1.5 mm, apex emarginate; filaments (1–)1.5–2 × 0.5 mm, anthers 0.4–0.6 mm; carpels 10–25; styles 1.8–2.2 mm. |
4–10 mm diam.; epicalyx bractlets linear to narrowly elliptic, 0.8–2(–3) × 0.2–0.4 mm, 1/2–2/3 length of sepals; hypanthium 1–2.5 × 2–5 mm, 1/2 to nearly as deep as wide, interior glabrous or pilose; sepals moderately reflexed, abaxially green to reddish, 1.5–3 mm; petals white, sometimes tinged with pink, linear to broadly oblanceolate, (1–)1.5–4(–4.5) × 0.3–1.5 mm, apex acute to rounded or mucronate; filaments 0.5–2 × 0.1–0.4(–0.8) mm, anthers 0.2–0.5 mm; carpels 5–15; styles 1–2.5 mm. |
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Achenes | light brown, 1.5 mm, smooth or slightly rugose. |
light brown, 1.5–2.5 mm, rugose. |
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2n | = 28. |
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Horkelia tenuiloba |
Horkelia tridentata |
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Phenology | Flowering summer. | |||||
Habitat | Sandy soil, openings, in chaparral, oak woodlands | |||||
Elevation | 50–500 m (200–1600 ft) | |||||
Distribution |
CA
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CA; OR
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Discussion | Of conservation concern. Horkelia tenuiloba occurs on the western edges of the northern Coast Ranges in Marin, Mendocino, and Sonoma counties. Populations from San Luis Obispo formerly included in this species now are part of H. yadonii. A specimen (M. K. C[urran], July 5, 1885, UC) unequivocally of H. tenuiloba purportedly from San Luis Obispo is in all likelihood mislabeled with respect to locality. Horkelia tenuiloba is commonly associated with seral openings in chaparral and woodlands and might be dependent on periodic disturbance by fire. W. L. Jepson (1909–1943, vol. 2) used Potentilla stenoloba (1895) for the species encompassing the types of Horkelia tenuiloba and P. micheneri. The epithet micheneri (1893) has priority at species rank within Potentilla, since P. tenuiloba (Torrey) Greene is a later homonym of P. tenuiloba Jordan. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Rydberg established Horkelia flavescens on the basis of its more congested inflorescence, purportedly yellowish petals, and broader filaments. As noted by D. D. Keck (1938), the petals are usually white, at least when fresh, and the other characters are highly variable; he reduced H. flavescens to a subspecies of H. tridentata, which is here treated at the rank of variety. In general, var. flavescens tends to be more decumbent with a single, loosely congested flower cluster and broader petals; these characters completely overlap and can vary independently. The presence or absence of hairs on the hypanthium interior is the single most reliably diagnostic character and is the primary determiner of the geographic distributions presented here. The exception is a cluster of populations from the extreme southern end of the coastal range of var. flavescens in Colusa, Lake, and Mendocino counties, California, which have glabrous hypanthia but otherwise have all the diagnostic characters of var. flavescens. Although provisionally included here within var. flavescens, these populations may be better treated as a separate variety. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 252. | FNA vol. 9, p. 269. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | H. fusca var. tenuiloba, Potentilla micheneri, P. stenoloba | |||||
Name authority | (Torrey) A. Gray: Proc. Amer. Acad. Arts 6: 529. (1865) | Torrey: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 84, plate 6. (1857) | ||||
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