Horkelia tenuiloba |
Horkelia daucifolia |
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Santa Rosa, Santa Rosa horkelia, Sonoma horkelia, thin-lobed, thin-lobed horkelia |
carrot leafed horkelia, carrot-leaf horkelia |
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Habit | Plants loosely matted, green. | Plants rosette-forming to tufted, rarely ± matted, grayish or green. | ||||||||
Stems | ascending to erect, 1–4 dm, hairs ± spreading. |
ascending to erect, 1.5–3.5 dm, hairs 2–3 mm proximally, glands sparse to dense distally. |
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Basal leaves | weakly planar to ± cylindric, 5–15(–20) × 0.5–1.5 cm; stipules entire; leaflets 8–16(–20) per side, ± overlapping especially distally, cuneate to flabellate, 3–10 × 2–10 mm, 1/2 to nearly as wide as long, divided 1/2–3/4+ to midrib into 3–8 linear to narrowly oblanceolate or narrowly elliptic lobes, sparsely villous at least marginally, often with a tuft of hairs apically. |
weakly planar to loosely cylindric, (4–)5–12(–17) × 0.8–3(–4) cm, usually sericeous to villous, sometimes glabrate; stipules pinnately divided into 3–7 linear to filiform lobes; leaflets 5–10 per side, ± overlapping, obovate to broadly obcordate, 5–15(–25) × 5–20 mm, ± as wide as long, divided 3/4+ to midrib into (0–)2–15 linear to oblanceolate lobes 0.4–2(–3) mm wide, these not restricted to apex. |
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Cauline leaves | 2–5. |
3–6; stipules 8–15 mm, deeply 3–7-lobed proximally, 1–3-lobed distally. |
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Inflorescences | open to congested, flowers arranged individually and in glomerules, these sometimes subcapitate. |
open to ± congested, flowers arranged individually or in ± corymbiform clusters. |
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Pedicels | 1–6 mm. |
2–10(–20) mm. |
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Flowers | 10 mm diam.; epicalyx bractlets narrowly to broadly lanceolate, 1–3 × 0.5–1 mm, slightly shorter than sepals, entire; hypanthium 1–1.2 × 2.5–4.5 mm, less than 1/2 as deep as wide, interior pilose; sepals spreading to reflexed, lanceolate, 3–5 mm; petals oblanceolate, 2.5–4.5 × 1.5 mm, apex emarginate; filaments (1–)1.5–2 × 0.5 mm, anthers 0.4–0.6 mm; carpels 10–25; styles 1.8–2.2 mm. |
10–15 mm diam.; epicalyx bractlets linear to linear-lanceolate, 2.5–3.5 × 0.5 mm, 2/3 to ± equal to length of sepals; hypanthium 1–1.5 × 3.5–5 mm, less than 1/2 as deep as wide, interior glabrous; sepals ± spreading, abaxially green, 3.5–7.5 mm; petals white to cream, often drying yellowish, cuneate to obcordate, (3.5–)4–8 × 2–8 mm, apex ± emarginate, sometimes rounded; filaments 1–3 × 0.5–1.5 mm, anthers 0.5–0.8 mm; carpels 5–15, styles 2–4 mm. |
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Achenes | light brown, 1.5 mm, smooth or slightly rugose. |
dark brown, 2.4–3 mm, smooth to ± roughened. |
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2n | = 28. |
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Horkelia tenuiloba |
Horkelia daucifolia |
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Phenology | Flowering summer. | |||||||||
Habitat | Sandy soil, openings, in chaparral, oak woodlands | |||||||||
Elevation | 50–500 m (200–1600 ft) | |||||||||
Distribution |
CA
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CA; OR
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Discussion | Of conservation concern. Horkelia tenuiloba occurs on the western edges of the northern Coast Ranges in Marin, Mendocino, and Sonoma counties. Populations from San Luis Obispo formerly included in this species now are part of H. yadonii. A specimen (M. K. C[urran], July 5, 1885, UC) unequivocally of H. tenuiloba purportedly from San Luis Obispo is in all likelihood mislabeled with respect to locality. Horkelia tenuiloba is commonly associated with seral openings in chaparral and woodlands and might be dependent on periodic disturbance by fire. W. L. Jepson (1909–1943, vol. 2) used Potentilla stenoloba (1895) for the species encompassing the types of Horkelia tenuiloba and P. micheneri. The epithet micheneri (1893) has priority at species rank within Potentilla, since P. tenuiloba (Torrey) Greene is a later homonym of P. tenuiloba Jordan. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). D. D. Keck (1938) decided that the leaflet characters previously used to recognize segregates of Horkelia daucifolia were random variables that have not become localized into geographic subunits. At the same time, he established subsp. latior D. D. Keck to accommodate a single collection with leaves approaching those of H. sericata, with the speculation that the population was an amphiploid derivative of the two species. Subsequent collections from the same general area (Scott Mountain, Siskiyou County, California) provide a full range of variation between the extreme with exceptionally wide leaflet lobes and the typical form occurring at the base of the mountain. This variant of the species is accordingly not recognized here, but the extremes with narrower, more numerous leaflet lobes are recognized as varieties that coincide with major river drainages (B. Ertter and J. L. Reveal 2007). In the descriptions below, the pedicels of all varieties are sparsely pilose in addition to being puberulent or not. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 252. | FNA vol. 9, p. 266. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae | ||||||||
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Subordinate taxa | ||||||||||
Synonyms | H. fusca var. tenuiloba, Potentilla micheneri, P. stenoloba | Potentilla daucifolia | ||||||||
Name authority | (Torrey) A. Gray: Proc. Amer. Acad. Arts 6: 529. (1865) | (Greene) Rydberg: Bull. Torrey Bot. Club 25: 55. (1898) | ||||||||
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