Horkelia tenuiloba |
Horkelia californica |
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Santa Rosa, Santa Rosa horkelia, Sonoma horkelia, thin-lobed, thin-lobed horkelia |
California horkelia |
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Habit | Plants loosely matted, green. | Plants ± tufted, green. | ||||||||
Stems | ascending to erect, 1–4 dm, hairs ± spreading. |
ascending to erect, (1–)5–10(–12) dm, hairs ± spreading. |
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Basal leaves | weakly planar to ± cylindric, 5–15(–20) × 0.5–1.5 cm; stipules entire; leaflets 8–16(–20) per side, ± overlapping especially distally, cuneate to flabellate, 3–10 × 2–10 mm, 1/2 to nearly as wide as long, divided 1/2–3/4+ to midrib into 3–8 linear to narrowly oblanceolate or narrowly elliptic lobes, sparsely villous at least marginally, often with a tuft of hairs apically. |
planar, (5–)8–40(–50) × 1.5–11 cm; stipules entire; leaflets 3–9 per side, separate to slightly overlapping, ovate to round or broadly elliptic, 5–60 × (7–)10–30(–40) mm, 1/2 to as wide as long, unlobed or irregularly cleft 1/6–3/4+ to midrib into 5–15 linear or oblanceolate to obovate coarsely toothed lobes, collectively 10–60-toothed, pilose to villous. |
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Cauline leaves | 2–5. |
4–10. |
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Inflorescences | open to congested, flowers arranged individually and in glomerules, these sometimes subcapitate. |
± open, flowers arranged individually and in glomerules. |
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Pedicels | 1–6 mm. |
1–20 mm. |
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Flowers | 10 mm diam.; epicalyx bractlets narrowly to broadly lanceolate, 1–3 × 0.5–1 mm, slightly shorter than sepals, entire; hypanthium 1–1.2 × 2.5–4.5 mm, less than 1/2 as deep as wide, interior pilose; sepals spreading to reflexed, lanceolate, 3–5 mm; petals oblanceolate, 2.5–4.5 × 1.5 mm, apex emarginate; filaments (1–)1.5–2 × 0.5 mm, anthers 0.4–0.6 mm; carpels 10–25; styles 1.8–2.2 mm. |
8–15 mm diam.; epicalyx bractlets narrowly oblong to ovate, 4–6(–8) × 1–2.5 mm, ± equal to sepals, entire or toothed; hypanthium 3–5.5 × 4–10 mm, 1/2 to nearly as deep as wide, interior glabrous or pilose; sepals spreading, lanceolate, 4–6.5(–8) mm; petals oblanceolate to elliptic or oblong, 3–8 × 1–4 mm, apex obtuse to rounded; filaments 0.5–3 × 0.2–1.5 mm, anthers (0.8–)1.3–1.8 mm; carpels (50–)80–200(–220); styles 2–4 mm. |
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Achenes | light brown, 1.5 mm, smooth or slightly rugose. |
brown, 0.8–1 mm, smooth or slightly rugose. |
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2n | = 28. |
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Horkelia tenuiloba |
Horkelia californica |
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Phenology | Flowering summer. | |||||||||
Habitat | Sandy soil, openings, in chaparral, oak woodlands | |||||||||
Elevation | 50–500 m (200–1600 ft) | |||||||||
Distribution |
CA
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CA
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Discussion | Of conservation concern. Horkelia tenuiloba occurs on the western edges of the northern Coast Ranges in Marin, Mendocino, and Sonoma counties. Populations from San Luis Obispo formerly included in this species now are part of H. yadonii. A specimen (M. K. C[urran], July 5, 1885, UC) unequivocally of H. tenuiloba purportedly from San Luis Obispo is in all likelihood mislabeled with respect to locality. Horkelia tenuiloba is commonly associated with seral openings in chaparral and woodlands and might be dependent on periodic disturbance by fire. W. L. Jepson (1909–1943, vol. 2) used Potentilla stenoloba (1895) for the species encompassing the types of Horkelia tenuiloba and P. micheneri. The epithet micheneri (1893) has priority at species rank within Potentilla, since P. tenuiloba (Torrey) Greene is a later homonym of P. tenuiloba Jordan. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Plants of Horkelia californica have the largest hypanthia in the genus, and the hypanthia are significantly deeper than others that are equally wide. Other unique features include the often toothed bractlets and the slightly clawed petals that are often inwardly curved with undulate margins. These features also help to distinguish the species from Drymocallis glandulosa, with which it is often confused: both species have a similar gross morphology and overlapping ranges and habitats. The three varieties recognized here were treated as distinct species by D. D. Keck (1938). The existence of intermediate populations makes varietal status the preferred option. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 252. | FNA vol. 9, p. 256. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | H. fusca var. tenuiloba, Potentilla micheneri, P. stenoloba | Potentilla californica | ||||||||
Name authority | (Torrey) A. Gray: Proc. Amer. Acad. Arts 6: 529. (1865) | Chamisso & Schlechtendal: Linnaea 2: 26. (1827) | ||||||||
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