Horkelia sericata |
Horkelia yadonii |
|
---|---|---|
Howell's horkelia, silky horkelia |
Santa Lucia horkelia, Yadon's or Santa Lucia horkelia |
|
Habit | Plants ± tufted, silvery. | Plants tufted to matted, ± grayish green. |
Stems | ascending to erect, 1.5–4.5 dm, hairs 1 mm proximally, glands absent or sparse distally. |
ascending to erect, (0.5–)2–6(–7) dm, hairs spreading. |
Basal leaves | ± cylindric to weakly planar, 3–10 × 0.3–1.2 cm, densely sericeous, often villous on margins apically; stipules usually entire or forked, rarely pinnately divided into linear lobes; leaflets (8–)10–20 per side, ± overlapping, elliptic to flabellate, 2–8 × 1–4 mm, 1/2–2/3 as wide as long, divided ± 1/2+ to midrib into (0–)2–4 elliptic lobes 1–2 mm wide, these not restricted to apex. |
± planar, (3–)6–20(–32) × 0.5–1.5(–2) cm; stipules entire; leaflets 7–16 per side, separate to ± overlapping especially distally, broadly cuneate to nearly round, (3–)4–15(–17) × (2–)5–12 mm, 3/4 to nearly as wide as long, divided 1/5–1/2 to midrib into (3–)8–15(–30) acute to obtuse teeth, often medially notched as well 1/3–2/3 to midrib, villous. |
Cauline leaves | (2 or)3–5; stipules 3–8 mm, entire or shallowly 1–2-toothed. |
3–5. |
Inflorescences | open, flowers arranged individually and/or in non-capitate glomerules. |
open, flowers arranged individually and in glomerules. |
Pedicels | 1–4 mm. |
1–7(–20) mm. |
Flowers | 10 mm diam.; epicalyx bractlets linear-lanceolate, 1.5–3 × 0.5 mm, ± 2/3 length of sepals; hypanthium 1–1.5 × 2–3 mm, ± 1/2 to as deep as wide, interior glabrous; sepals spreading to reflexed, abaxially green to reddish or purplish, 2–4 mm; petals white to pink or red-veined, narrowly obcordate, 3–4.5(–7) × 2–3 mm, apex ± emarginate; filaments 0.5–1.5 × 0.2–0.5 mm, anthers 0.4–0.6 mm; carpels 2–6; styles 1.5–2 mm. |
8–12 mm diam.; epicalyx bractlets lanceolate to ovate, 1.5–4.5 × 1–2(–2.5) mm, 1/2–3/4 length of sepals, entire; hypanthium 1.5–3 × 3–6.5 mm, less than 1/2 as deep as wide, interior sparsely pilose; sepals spreading to ± reflexed, lanceolate, (3–)4–6.5 mm; petals broadly oblanceolate-elliptic, 3–5.5 × 2–3 mm, apex truncate; filaments 1–2 × 0.5–1 mm, anthers 0.8–1.2 mm; carpels 20–60; styles 2.5–3 mm. |
Achenes | brown, 2–2.5 mm, smooth. |
brown, 1.5 mm, smooth. |
2n | = 28. |
|
Horkelia sericata |
Horkelia yadonii |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Chaparral, oak-conifer woodlands, on serpentine-derived soil | Granitic sand, meadows, along stream banks, in chaparral, conifer woodlands |
Elevation | 100–1200 m (300–3900 ft) | 300–1900 m (1000–6200 ft) |
Distribution |
CA; OR
|
CA |
Discussion | Of conservation concern. D. D. Keck (1938) expanded the circumscription of Horkelia sericata to encompass the plants segregated here as H. howellii, on the grounds that intergradation is too extensive to justify taxonomic recognition of the extremes. In this evaluation, however, most collections can be unequivocally divided between plants with compact silvery-sericeous leaves with entire or forked stipules (H. sericata), and plants with larger, greener leaves and pinnately divided stipules (H. howellii). It is not known if the chromosome count provided by P. A. Munz (1959) was derived from H. howellii or H. sericata. As here circumscribed, Horkelia sericata is a localized taxon known only from Curry County, Oregon, and the Gasquet serpentine area in adjacent Del Norte County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Horkelia yadonii is known only from localized populations in the La Panza, San Rafael, and Santa Lucia ranges of Monterey, San Luis Obispo, and Santa Barbara counties. The isolated populations often differ in such characters as general size, leaf proportions, and shape of epicalyx bractlets. The most distinctive extreme, in Monterey County, is characterized by relatively long, narrow leaves, small, few-toothed leaflets, long pedicels, few small flowers, and ovate bractlets. These differences, however, are not consistently or sharply defined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 265. | FNA vol. 9, p. 253. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia |
Sibling taxa | ||
Synonyms | Potentilla sericata | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 20: 364. (1885) | Ertter: Syst. Bot. 18: 139, fig. 2. (1993) |
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