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Howell's horkelia, silky horkelia

Santa Lucia horkelia, Yadon's or Santa Lucia horkelia

Habit Plants ± tufted, silvery. Plants tufted to matted, ± grayish green.
Stems

ascending to erect, 1.5–4.5 dm, hairs 1 mm proximally, glands absent or sparse distally.

ascending to erect, (0.5–)2–6(–7) dm, hairs spreading.

Basal leaves

± cylindric to weakly planar, 3–10 × 0.3–1.2 cm, densely sericeous, often villous on margins apically;

stipules usually entire or forked, rarely pinnately divided into linear lobes;

leaflets (8–)10–20 per side, ± overlapping, elliptic to flabellate, 2–8 × 1–4 mm, 1/2–2/3 as wide as long, divided ± 1/2+ to midrib into (0–)2–4 elliptic lobes 1–2 mm wide, these not restricted to apex.

± planar, (3–)6–20(–32) × 0.5–1.5(–2) cm;

stipules entire;

leaflets 7–16 per side, separate to ± overlapping especially distally, broadly cuneate to nearly round, (3–)4–15(–17) × (2–)5–12 mm, 3/4 to nearly as wide as long, divided 1/5–1/2 to midrib into (3–)8–15(–30) acute to obtuse teeth, often medially notched as well 1/3–2/3 to midrib, villous.

Cauline leaves

(2 or)3–5;

stipules 3–8 mm, entire or shallowly 1–2-toothed.

3–5.

Inflorescences

open, flowers arranged individually and/or in non-capitate glomerules.

open, flowers arranged individually and in glomerules.

Pedicels

1–4 mm.

1–7(–20) mm.

Flowers

10 mm diam.;

epicalyx bractlets linear-lanceolate, 1.5–3 × 0.5 mm, ± 2/3 length of sepals;

hypanthium 1–1.5 × 2–3 mm, ± 1/2 to as deep as wide, interior glabrous;

sepals spreading to reflexed, abaxially green to reddish or purplish, 2–4 mm;

petals white to pink or red-veined, narrowly obcordate, 3–4.5(–7) × 2–3 mm, apex ± emarginate;

filaments 0.5–1.5 × 0.2–0.5 mm, anthers 0.4–0.6 mm;

carpels 2–6;

styles 1.5–2 mm.

8–12 mm diam.;

epicalyx bractlets lanceolate to ovate, 1.5–4.5 × 1–2(–2.5) mm, 1/2–3/4 length of sepals, entire;

hypanthium 1.5–3 × 3–6.5 mm, less than 1/2 as deep as wide, interior sparsely pilose;

sepals spreading to ± reflexed, lanceolate, (3–)4–6.5 mm;

petals broadly oblanceolate-elliptic, 3–5.5 × 2–3 mm, apex truncate;

filaments 1–2 × 0.5–1 mm, anthers 0.8–1.2 mm;

carpels 20–60;

styles 2.5–3 mm.

Achenes

brown, 2–2.5 mm, smooth.

brown, 1.5 mm, smooth.

2n

= 28.

Horkelia sericata

Horkelia yadonii

Phenology Flowering summer. Flowering summer.
Habitat Chaparral, oak-conifer woodlands, on serpentine-derived soil Granitic sand, meadows, along stream banks, in chaparral, conifer woodlands
Elevation 100–1200 m (300–3900 ft) 300–1900 m (1000–6200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Of conservation concern.

D. D. Keck (1938) expanded the circumscription of Horkelia sericata to encompass the plants segregated here as H. howellii, on the grounds that intergradation is too extensive to justify taxonomic recognition of the extremes. In this evaluation, however, most collections can be unequivocally divided between plants with compact silvery-sericeous leaves with entire or forked stipules (H. sericata), and plants with larger, greener leaves and pinnately divided stipules (H. howellii). It is not known if the chromosome count provided by P. A. Munz (1959) was derived from H. howellii or H. sericata.

As here circumscribed, Horkelia sericata is a localized taxon known only from Curry County, Oregon, and the Gasquet serpentine area in adjacent Del Norte County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Horkelia yadonii is known only from localized populations in the La Panza, San Rafael, and Santa Lucia ranges of Monterey, San Luis Obispo, and Santa Barbara counties. The isolated populations often differ in such characters as general size, leaf proportions, and shape of epicalyx bractlets. The most distinctive extreme, in Monterey County, is characterized by relatively long, narrow leaves, small, few-toothed leaflets, long pedicels, few small flowers, and ovate bractlets. These differences, however, are not consistently or sharply defined.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 265. FNA vol. 9, p. 253.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae
Synonyms Potentilla sericata
Name authority S. Watson: Proc. Amer. Acad. Arts 20: 364. (1885) Ertter: Syst. Bot. 18: 139, fig. 2. (1993)
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