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Howell's horkelia, silky horkelia

Kern Plateau horkelia, Tulare cinquefoil

Habit Plants ± tufted, silvery. Plants 1–4 dm diam.
Stems

ascending to erect, 1.5–4.5 dm, hairs 1 mm proximally, glands absent or sparse distally.

decumbent to erect, 0.3–1.8(–2.5) dm.

Basal leaves

± cylindric to weakly planar, 3–10 × 0.3–1.2 cm, densely sericeous, often villous on margins apically;

stipules usually entire or forked, rarely pinnately divided into linear lobes;

leaflets (8–)10–20 per side, ± overlapping, elliptic to flabellate, 2–8 × 1–4 mm, 1/2–2/3 as wide as long, divided ± 1/2+ to midrib into (0–)2–4 elliptic lobes 1–2 mm wide, these not restricted to apex.

2–8(–10) × 0.5–1.3 cm;

leaflets 4–6(–10) per side, overlapping, cuneate to flabellate, 3–8 mm, divided 3/4+ to midrib into (3–)5–8 oblong to obovate or oblanceolate lobes, villous to pilose at least marginally or apically.

Cauline leaves

(2 or)3–5;

stipules 3–8 mm, entire or shallowly 1–2-toothed.

1 or 2.

Inflorescences

open, flowers arranged individually and/or in non-capitate glomerules.

Pedicels

1–4 mm.

2–7 mm.

Flowers

10 mm diam.;

epicalyx bractlets linear-lanceolate, 1.5–3 × 0.5 mm, ± 2/3 length of sepals;

hypanthium 1–1.5 × 2–3 mm, ± 1/2 to as deep as wide, interior glabrous;

sepals spreading to reflexed, abaxially green to reddish or purplish, 2–4 mm;

petals white to pink or red-veined, narrowly obcordate, 3–4.5(–7) × 2–3 mm, apex ± emarginate;

filaments 0.5–1.5 × 0.2–0.5 mm, anthers 0.4–0.6 mm;

carpels 2–6;

styles 1.5–2 mm.

3–15, 8–10 mm diam.;

epicalyx bractlets linear-lanceolate, 1–2.5(–3) × 0.3–0.5(–0.8) mm, ± 2/3 length of sepals;

hypanthium 1–1.5 × 2.5–4.5 mm, nearly 1/2 as deep as wide, interior sparsely pilose;

sepals spreading to reflexed, broadly lanceolate, (2–)3–4.5(–5) mm, hairs stiff, 0.5–1 mm;

petals not pink-tinged, narrowly oblanceolate to narrowly oblong, 2–4 mm, apex rounded to truncate, often slightly mucronate;

filaments white, 1–2 × 0.3–0.5 mm, usually glabrous, rarely sparsely pilose, anthers 0.6–0.8 mm;

carpels 5–12;

styles 1.5–2 mm.

Achenes

brown, 2–2.5 mm, smooth.

light brown to brown, 2–2.5 mm.

2n

= 28.

Horkelia sericata

Horkelia tularensis

Phenology Flowering summer. Flowering summer.
Habitat Chaparral, oak-conifer woodlands, on serpentine-derived soil Dry, rocky metamorphic ridges, in subalpine conifer woodlands
Elevation 100–1200 m (300–3900 ft) 2300–2900 m (7500–9500 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Of conservation concern.

D. D. Keck (1938) expanded the circumscription of Horkelia sericata to encompass the plants segregated here as H. howellii, on the grounds that intergradation is too extensive to justify taxonomic recognition of the extremes. In this evaluation, however, most collections can be unequivocally divided between plants with compact silvery-sericeous leaves with entire or forked stipules (H. sericata), and plants with larger, greener leaves and pinnately divided stipules (H. howellii). It is not known if the chromosome count provided by P. A. Munz (1959) was derived from H. howellii or H. sericata.

As here circumscribed, Horkelia sericata is a localized taxon known only from Curry County, Oregon, and the Gasquet serpentine area in adjacent Del Norte County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Horkelia tularensis is known only from the Kern Plateau in Tulare County. Greenish and reddish plants can grow intermixed in a single population.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 265. FNA vol. 9, p. 249.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Hispidulae
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. wilderae, H. yadonii
Synonyms Potentilla sericata Potentilla tularensis
Name authority S. Watson: Proc. Amer. Acad. Arts 20: 364. (1885) (J. T. Howell) Munz: Suppl. Calif. Fl., 110. (1968)
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