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Howell's horkelia, silky horkelia

White Mountain horkelia, White Mountains. horkelia

Habit Plants ± tufted, silvery. Plants 0.7–3 dm diam.
Stems

ascending to erect, 1.5–4.5 dm, hairs 1 mm proximally, glands absent or sparse distally.

ascending to erect, 1–2.5 dm.

Basal leaves

± cylindric to weakly planar, 3–10 × 0.3–1.2 cm, densely sericeous, often villous on margins apically;

stipules usually entire or forked, rarely pinnately divided into linear lobes;

leaflets (8–)10–20 per side, ± overlapping, elliptic to flabellate, 2–8 × 1–4 mm, 1/2–2/3 as wide as long, divided ± 1/2+ to midrib into (0–)2–4 elliptic lobes 1–2 mm wide, these not restricted to apex.

3–10 × 0.4–0.8 cm;

leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid.

Cauline leaves

(2 or)3–5;

stipules 3–8 mm, entire or shallowly 1–2-toothed.

3–7.

Inflorescences

open, flowers arranged individually and/or in non-capitate glomerules.

Pedicels

1–4 mm.

2–8(–12) mm.

Flowers

10 mm diam.;

epicalyx bractlets linear-lanceolate, 1.5–3 × 0.5 mm, ± 2/3 length of sepals;

hypanthium 1–1.5 × 2–3 mm, ± 1/2 to as deep as wide, interior glabrous;

sepals spreading to reflexed, abaxially green to reddish or purplish, 2–4 mm;

petals white to pink or red-veined, narrowly obcordate, 3–4.5(–7) × 2–3 mm, apex ± emarginate;

filaments 0.5–1.5 × 0.2–0.5 mm, anthers 0.4–0.6 mm;

carpels 2–6;

styles 1.5–2 mm.

3–15, 10 mm diam.;

epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals;

hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose;

sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm;

petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate;

filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm;

carpels (10–)12–18(–20);

styles 1.8–2.2 mm.

Achenes

brown, 2–2.5 mm, smooth.

brown to dark brown, 1.5–2 mm.

2n

= 28.

Horkelia sericata

Horkelia hispidula

Phenology Flowering summer. Flowering summer.
Habitat Chaparral, oak-conifer woodlands, on serpentine-derived soil Dry, rocky alpine flats, in subalpine conifer woodlands
Elevation 100–1200 m (300–3900 ft) 3000–3400 m (9800–11200 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Of conservation concern.

D. D. Keck (1938) expanded the circumscription of Horkelia sericata to encompass the plants segregated here as H. howellii, on the grounds that intergradation is too extensive to justify taxonomic recognition of the extremes. In this evaluation, however, most collections can be unequivocally divided between plants with compact silvery-sericeous leaves with entire or forked stipules (H. sericata), and plants with larger, greener leaves and pinnately divided stipules (H. howellii). It is not known if the chromosome count provided by P. A. Munz (1959) was derived from H. howellii or H. sericata.

As here circumscribed, Horkelia sericata is a localized taxon known only from Curry County, Oregon, and the Gasquet serpentine area in adjacent Del Norte County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 265. FNA vol. 9, p. 249.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Hispidulae
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Synonyms Potentilla sericata Potentilla hispidula
Name authority S. Watson: Proc. Amer. Acad. Arts 20: 364. (1885) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908)
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