Horkelia sect. Parryae |
Rosaceae tribe Potentilleae |
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Habit | Plants forming rosettes or open mats, green, conspicuously glandular, ± resinously aromatic. | Herbs, perennial, rarely annual or biennial, shrubs, or subshrubs; unarmed. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | prostrate to erect, (0.5–)1–3(–3.5) dm. |
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Leaves | alternate, rarely opposite, pinnately (palmately) compound (simple in Alchemilla, Aphanes, and Chamaerhodos); stipules persistent (absent in Chamaerhodos), adnate to petiole; venation pinnate or palmate. |
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Basal leaves | planar; stipules usually entire, rarely deeply 2-lobed; leaflets 3–7 per side, separate, divided ± 1/4–3/4 to midrib into 5–15 teeth or lobes not restricted to apex. |
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Inflorescences | open, flowers arranged individually. |
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Pedicels | becoming reflexed or recurved, 3–15 mm. |
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Flowers | epicalyx bractlets linear-lanceolate to elliptic or ovate, 0.3–1.5 mm wide, entire; hypanthium interior glabrous; sepals acute; petals white, elliptic or oblong to obovate, apex obtuse to rounded or slightly emarginate; filaments white, glabrous, anthers at least slightly longer than wide; carpels 3–4 or (17–)20–50. |
perianth and androecium perigynous; epicalyx bractlets present, sometimes absent; hypanthium usually patelliform, cupulate, or campanulate, sometimes turbinate, saucer-shaped, flat-bottomed, or subglobose to ellipsoid or ovoid; torus flat to conic or turbinate, enlarged (absent or reduced in Alchemilla, Aphanes, and Chamaerhodos); carpels 1–260, styles basal or lateral to subterminal, distinct; ovules 1(or 2), basal. |
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Fruits | aggregated achenes (achenes in Alchemilla and Aphanes); torus sometimes fleshy; styles deciduous or persistent, not elongate. |
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Achenes | 1.3–2.2 mm, finely reticulate or coarsely rugose. |
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Horkelia sect. Parryae |
Rosaceae tribe Potentilleae |
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Distribution |
CA |
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Pacific Islands; Australia |
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Discussion | Species 2 (2 in the flora). Section Parryae accommodates two localized species with open inflorescences in which the slender pedicels become reflexed to recurved at or soon after flowering. P. A. Rydberg (1908c) included both species in his group Cuneatae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 14–22, species ca. 860 (14 genera, 189 species, including 1 hybrid, in the flora area). The base chromosome number for Potentilleae is mostly x = 7 (8 in Alchemilla and Aphanes; 14 in Comarum). Variation in the number of genera recognized in Potentilleae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of Potentilla and segregates here (see 9. Ivesia and 8. Potentilla for discussion). In the former, Duchesnea, Horkelia, Horkeliella, and Ivesia are included within Potentilla. Likewise, Aphanes is included within Alchemilla by Potter et al. while it is kept distinct here. Potentilla and its segregates and Fragaria are host to Phragmidium rusts, but not the other genera of the tribe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 257. | FNA vol. 9, p. 119. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Ertter & Reveal: Novon 17: 316. (2007) | Sweet: Brit. Fl. Gard. 2: sub plate 124. (1825) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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