Horkelia sect. Hispidulae
Rosaceae subfam. rosoideae
decumbent to erect, 0.3–2.5 dm.
alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;
stipules present, rarely absent.
± cylindric to weakly planar;
stipules entire;
leaflets 4–14 per side, overlapping at least distally, divided 1/2–3/4+ to midrib into 3–8 lobes not restricted to apex.
± congested, flowers arranged in dense corymbiform clusters.
remaining straight, 2–8(–12) mm.
epicalyx bractlets linear to lanceolate, 0.3–0.5(–0.8) mm wide, entire;
hypanthium interior sparsely pilose to densely villous;
sepals acute;
petals white, sometimes pink-tinged, narrowly oblanceolate to narrowly elliptic or oblong, apex usually acute to rounded to truncate, sometimes slightly mucronate or emarginate;
filaments white to pinkish, glabrous or sparsely pilose adaxially, anthers longer than wide;
carpels 5–18(–20).
torus usually enlarged, sometimes small or absent;
carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);
ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).
achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;
styles persistent or deciduous, not elongate (elongate but not plumose in Geum).
1.5–2.5 mm, smooth.
= 7(8).
Horkelia sect. Hispidulae
Rosaceae subfam. rosoideae
Species 3 (3 in the flora).
Section Hispidulae accommodates three widely separated, localized species, with plants all forming tight, densely hairy mats in rocky sites at elevations of 2000 to 3400 m in the White Mountains, southern Sierra Nevada, and Siskiyou Mountains of California, adjacent Nevada, and southwestern Oregon.
When describing Horkelia tularensis (as Potentilla tularensis), Howell compared it to H. hispidula with the speculation that these interior species bordering the Great Basin were relicts of an evolutionary line distinct from the more coastal species. Given the geographic proximity to Horkeliella, which serves as a morphologic bridge between Horkelia and Ivesia, it is possible that sect. Hispidulae represents the relictual ancestral radiation within the genus.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.
Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Sepals narrowly lanceolate, with silky hairs 1–1.5 mm; hypanthium interior densely villous; petals often pink-tinged; Siskiyou Mountains, California and Oregon. | H. hendersonii |
1. Sepals broadly lanceolate, with stiff hairs 0.5–1 mm; hypanthium interior sparsely pilose; petals not pink-tinged; Sierra Nevada and White Mountains, California and Nevada | → 2 |
2. Basal leaves: leaflets 4–6(–10) per side; cauline leaves 1 or 2; carpels 5–12; s Sierra Nevada. | H. tularensis |
2. Basal leaves: leaflets (6–)10–14 per side; cauline leaves 3–7; carpels (10–)12–18(–20); White Mountains. | H. hispidula |
WildflowerSearch
iNaturalist
