Horkelia parryi |
Horkelia rydbergii |
|
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chamise horkelia, Parry horkelia, Parry's horkelia |
Rydberg's horkelia |
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Habit | Plants forming open mats. | Plants tufted to ± matted, usually grayish to grayish green. |
Stems | ascending to erect, 1–3(–3.5) dm. |
ascending to erect, (1–)2–7 dm, hairs ascending to appressed. |
Basal leaves | 4–10(–12) × 1–2 cm; leaflets 3–6(or 7) per side, narrowly obovate, 5–12(–15) × 4–8(–10) mm, 1/2–3/4 as wide as long, divided ± 1/4 to midrib into 5–10 oblong to broadly obovate teeth, sparsely pilose especially on midveins and margins. |
± planar, (4–)8–20(–30) × 0.8–2(–3) cm; stipules entire; leaflets 7–14 per side, separate to ± overlapping at least distally, cuneate to flabellate, (3–)5–15(–22) × 3–10(–15) mm, ± 1/2 to as wide as long, divided ± 1/3 to midrib into (3–)5–10 acute to obtuse teeth, densely (to sparsely) strigose or pilose. |
Cauline leaves | 1–4. |
(2 or)3–6(–8). |
Inflorescences | ± open to congested, flowers arranged individually or in glomerules. |
|
Pedicels | (3–)5–15 mm. |
(1–)2–8 mm. |
Flowers | 15 mm diam.; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm, ± 2/3 length of sepals; hypanthium 0.6–1 × 2.5–4 mm, less than 1/3 as deep as wide; sepals ± spreading to reflexed, lanceolate to narrowly elliptic, 3.5–6 mm; petals elliptic to obovate, 4–7 × 3 mm, apex obtuse to rounded to truncate or slightly emarginate; filaments 1–3 × 0.6–1.3 mm, anthers 0.6–1 mm; carpels (17–)20–50; styles 1.5–2.5 mm. |
10–15 mm diam.; epicalyx bractlets lanceolate to narrowly elliptic, 2–3.5(–4) × 0.5–1 mm, ± 3/4 length of sepals, entire; hypanthium 1–1.5(–2) × 2.5–4.5(–6) mm, less than 1/2 as deep as wide, interior pilose; sepals spreading to ± reflexed, lanceolate, (2.5–)3–5(–5.2) mm; petals oblong to oblanceolate, 4–5.5 × 1.2–2.5 mm, apex rounded to truncate or slightly emarginate; filaments 0.5–2 × 0.5–1 mm, anthers 0.6–1 mm; carpels 20–50(–120); styles 2–4 mm. |
Achenes | grayish, 1.3–1.5 mm, finely reticulate. |
light to dark brown, 1–1.5 mm, smooth or roughened. |
2n | = 28. |
|
Horkelia parryi |
Horkelia rydbergii |
|
Phenology | Flowering late spring–early summer. | Flowering summer. |
Habitat | Chaparral, pine-oak woodlands, primarily on Ione Formation, rarely on schist or limestone | Dry to moist meadows and stream banks, in conifer woodlands |
Elevation | 80–900 m (300–3000 ft) | 1200–2800 m (3900–9200 ft) |
Distribution |
CA
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CA
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Discussion | Of conservation concern. Horkelia parryi is known from the foothills of the western Sierra Nevada in Amador, Calaveras, and El Dorado counties, and is disjunct in Mariposa County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Horkelia rydbergii occurs in the Transverse Ranges of Kern, Los Angeles, San Bernardino, and Ventura counties. Reports from farther north (as H. bolanderi var. parryi) are referable to H. marinensis, H. yadonii, or, possibly, H. cuneata. Although there has been occasional nomenclatural confusion between this taxon (as H. bolanderi var. parryi) and H. parryi Greene, the two are taxonomically and geographically distinct. Plants in the Mount Pinos–Lockwood Valley area of Kern and Ventura counties, and on Frazier Mountain, are distinctly gray with an abundance of tightly curled hairs. The leaves tend to be relatively short (to 8 cm) with relatively small leaflets (3–6 mm) on the basal leaves. Stems are relatively short (to 3 dm), and the fruits are dark brown, distinctly rugose, and 1.3–1.5 mm. This phase (including the type of Horkelia rydbergii) abruptly gives way to a grayish green phase with a slightly less dense, pilose indument in the San Bernardino and San Gabriel mountains. Here the plants tend to have larger basal leaves (to 30 cm) and leaflets (5–12 mm), longer stems (to 7 dm), and fruits that are a lighter brown, mostly smooth, and 1–1.2 mm. This phase in turn grades into even less densely hairy plants near Bear Lake, where the type of H. bolanderi var. parryi was obtained. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 258. | FNA vol. 9, p. 251. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Potentilla parryi | H. bernardina, H. bolanderi subsp. parryi, H. bolanderi var. parryi, Potentilla bolanderi var. parryi |
Name authority | Greene: Bull. Calif. Acad. Sci. 2: 416. (1887) | Elmer: Bot. Gaz. 39: 50. (1905) |
Web links |