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chamise horkelia, Parry horkelia, Parry's horkelia

carrot leafed horkelia, carrot-leaf horkelia

Habit Plants forming open mats. Plants rosette-forming to tufted, rarely ± matted, grayish or green.
Stems

ascending to erect, 1–3(–3.5) dm.

ascending to erect, 1.5–3.5 dm, hairs 2–3 mm proximally, glands sparse to dense distally.

Basal leaves

4–10(–12) × 1–2 cm;

leaflets 3–6(or 7) per side, narrowly obovate, 5–12(–15) × 4–8(–10) mm, 1/2–3/4 as wide as long, divided ± 1/4 to midrib into 5–10 oblong to broadly obovate teeth, sparsely pilose especially on midveins and margins.

weakly planar to loosely cylindric, (4–)5–12(–17) × 0.8–3(–4) cm, usually sericeous to villous, sometimes glabrate;

stipules pinnately divided into 3–7 linear to filiform lobes;

leaflets 5–10 per side, ± overlapping, obovate to broadly obcordate, 5–15(–25) × 5–20 mm, ± as wide as long, divided 3/4+ to midrib into (0–)2–15 linear to oblanceolate lobes 0.4–2(–3) mm wide, these not restricted to apex.

Cauline leaves

1–4.

3–6;

stipules 8–15 mm, deeply 3–7-lobed proximally, 1–3-lobed distally.

Inflorescences

open to ± congested, flowers arranged individually or in ± corymbiform clusters.

Pedicels

(3–)5–15 mm.

2–10(–20) mm.

Flowers

15 mm diam.;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm, ± 2/3 length of sepals;

hypanthium 0.6–1 × 2.5–4 mm, less than 1/3 as deep as wide;

sepals ± spreading to reflexed, lanceolate to narrowly elliptic, 3.5–6 mm;

petals elliptic to obovate, 4–7 × 3 mm, apex obtuse to rounded to truncate or slightly emarginate;

filaments 1–3 × 0.6–1.3 mm, anthers 0.6–1 mm;

carpels (17–)20–50;

styles 1.5–2.5 mm.

10–15 mm diam.;

epicalyx bractlets linear to linear-lanceolate, 2.5–3.5 × 0.5 mm, 2/3 to ± equal to length of sepals;

hypanthium 1–1.5 × 3.5–5 mm, less than 1/2 as deep as wide, interior glabrous;

sepals ± spreading, abaxially green, 3.5–7.5 mm;

petals white to cream, often drying yellowish, cuneate to obcordate, (3.5–)4–8 × 2–8 mm, apex ± emarginate, sometimes rounded;

filaments 1–3 × 0.5–1.5 mm, anthers 0.5–0.8 mm;

carpels 5–15, styles 2–4 mm.

Achenes

grayish, 1.3–1.5 mm, finely reticulate.

dark brown, 2.4–3 mm, smooth to ± roughened.

2n

= 28.

Horkelia parryi

Horkelia daucifolia

Phenology Flowering late spring–early summer.
Habitat Chaparral, pine-oak woodlands, primarily on Ione Formation, rarely on schist or limestone
Elevation 80–900 m (300–3000 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Horkelia parryi is known from the foothills of the western Sierra Nevada in Amador, Calaveras, and El Dorado counties, and is disjunct in Mariposa County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

D. D. Keck (1938) decided that the leaflet characters previously used to recognize segregates of Horkelia daucifolia were random variables that have not become localized into geographic subunits. At the same time, he established subsp. latior D. D. Keck to accommodate a single collection with leaves approaching those of H. sericata, with the speculation that the population was an amphiploid derivative of the two species. Subsequent collections from the same general area (Scott Mountain, Siskiyou County, California) provide a full range of variation between the extreme with exceptionally wide leaflet lobes and the typical form occurring at the base of the mountain. This variant of the species is accordingly not recognized here, but the extremes with narrower, more numerous leaflet lobes are recognized as varieties that coincide with major river drainages (B. Ertter and J. L. Reveal 2007).

In the descriptions below, the pedicels of all varieties are sparsely pilose in addition to being puberulent or not.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet lobes 2–5(–8), linear to oblanceolate, 1–2(–3) mm wide; petals cuneate to obovate, 2–4 mm wide.
var. daucifolia
1. Leaflet lobes (2–)5–15, linear, 0.4–1 mm wide; petals broadly obovate to obcordate, 4–8 mm wide
→ 2
2. Pedicels 2–7(–10) mm, puberulent, eglandular; Oregon.
var. caruifolia
2. Pedicels 5–10(–20) mm, not puberulent, minutely glandular; California.
var. indicta
Source FNA vol. 9, p. 258. FNA vol. 9, p. 266.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Parryae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Subordinate taxa
H. daucifolia var. caruifolia, H. daucifolia var. daucifolia, H. daucifolia var. indicta
Synonyms Potentilla parryi Potentilla daucifolia
Name authority Greene: Bull. Calif. Acad. Sci. 2: 416. (1887) (Greene) Rydberg: Bull. Torrey Bot. Club 25: 55. (1898)
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