Horkelia parryi |
Horkelia congesta |
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chamise horkelia, Parry horkelia, Parry's horkelia |
shaggy horkelia, Sierra horkelia |
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Habit | Plants forming open mats. | Plants tufted or rosette-forming, green to grayish green. | ||||
Stems | ascending to erect, 1–3(–3.5) dm. |
ascending to erect, 1.5–5 dm, hairs 2–3 mm proximally, glands dense distally. |
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Basal leaves | 4–10(–12) × 1–2 cm; leaflets 3–6(or 7) per side, narrowly obovate, 5–12(–15) × 4–8(–10) mm, 1/2–3/4 as wide as long, divided ± 1/4 to midrib into 5–10 oblong to broadly obovate teeth, sparsely pilose especially on midveins and margins. |
± planar, (3–)4–15 × 1–3 cm, usually villous at least abaxially, often densely so; stipules entire, deeply 2-lobed, or pinnately divided into 3–5 linear lobes; leaflets 2–6 per side, ± separate, narrowly cuneate or lanceolate-elliptic to oblong, 5–20 × (1–)2–5(–6) mm, 1/5–1/2 as wide as long, divided 1/6–1/4 or less to midrib into (0–)3(–5) ± ovate teeth 1–2 mm wide, these restricted to apex. |
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Cauline leaves | 1–4. |
2–7; stipules 5–25(–30) mm, divided 3/4+ into linear or linear-lanceolate lobes, or incompletely divided or divided no more than 2/3 into lanceolate, often acuminate teeth. |
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Inflorescences | open to congested, flowers usually arranged in ± capitate glomerules or corymbiform clusters. |
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Pedicels | (3–)5–15 mm. |
1–3(–6) mm. |
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Flowers | 15 mm diam.; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm, ± 2/3 length of sepals; hypanthium 0.6–1 × 2.5–4 mm, less than 1/3 as deep as wide; sepals ± spreading to reflexed, lanceolate to narrowly elliptic, 3.5–6 mm; petals elliptic to obovate, 4–7 × 3 mm, apex obtuse to rounded to truncate or slightly emarginate; filaments 1–3 × 0.6–1.3 mm, anthers 0.6–1 mm; carpels (17–)20–50; styles 1.5–2.5 mm. |
8–16 mm diam.; epicalyx bractlets linear to lanceolate or narrowly elliptic, 1–3 × 0.3–0.8 mm, ± 2/3 length of sepals; hypanthium 2–2.5 × 3–4.5 mm, ± 1/2 to nearly as deep as wide, interior usually glabrous, sometimes sparsely pilose; sepals spreading, abaxially green, 2–4.5 mm; petals white to cream, ± obovate, (2.5–)3–6 × (1.5–)2–5 mm, apex ± rounded to ± emarginate; filaments (0.5–)1–2 × (0.2–)0.4–0.6 mm, anthers 0.4–0.7(–1) mm; carpels (6–)10–15(–20), styles 1.5–3 mm. |
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Achenes | grayish, 1.3–1.5 mm, finely reticulate. |
brown, 1.6–2.2 mm, faintly rugose. |
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2n | = 28. |
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Horkelia parryi |
Horkelia congesta |
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Phenology | Flowering late spring–early summer. | |||||
Habitat | Chaparral, pine-oak woodlands, primarily on Ione Formation, rarely on schist or limestone | |||||
Elevation | 80–900 m (300–3000 ft) | |||||
Distribution |
CA
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CA; OR
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Discussion | Of conservation concern. Horkelia parryi is known from the foothills of the western Sierra Nevada in Amador, Calaveras, and El Dorado counties, and is disjunct in Mariposa County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). As discussed by D. D. Keck (1938) and analyzed by T. N. Kaye (unpubl., Oregon Dept. of Agriculture, 1995), Horkelia congesta comprises three population clusters that form a latitudinal gradient in Oregon west of the Cascade Range, transitioning to H. tridentata in the mountains of California. Populations of H. congesta in remnant prairies in the Willamette Valley (probably including the type of H. hirsuta Lindley) have relatively tall, erect stems, pectinate stipules, relatively numerous, narrow leaflets, openly branched inflorescences with flowers in corymbiform clusters, and clawed petals 5–6 × 4–5 mm. In contrast, populations centered in rocky serpentine flats within the Illinois River drainage in Josephine County, which were described by Keck as subsp. nemorosa, tend to have shorter ascending stems, less divided stipules, fewer and broader leaflets, more capitate inflorescences, and scarcely clawed petals 3–4 × 2–3 mm. The differences between these two extremes are blurred by populations from the Umpqua Valley in Douglas County (the probable type locality of H. congesta), as well as scattered populations in northern Jackson and Josephine counties. Based on morphometric analyses of 19 characters, Kaye concluded that the Umpqua material overlapped slightly with plants from the Willamette Valley, while the Josephine County populations clustered separately. He retained the Umpqua and Willamette valley populations as subsp. congesta, with mid stem stipule features of greatest diagnostic value. This taxonomic conclusion is adopted here, with a change of rank from subspecies to variety. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 258. | FNA vol. 9, p. 267. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Potentilla parryi | Potentilla congesta | ||||
Name authority | Greene: Bull. Calif. Acad. Sci. 2: 416. (1887) | Douglas ex Hooker: Bot. Mag. 56: plate 2880. (1829) | ||||
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