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chamise horkelia, Parry horkelia, Parry's horkelia

Bolander's horkelia, border horkelia

Habit Plants forming open mats. Plants openly matted, grayish.
Stems

ascending to erect, 1–3(–3.5) dm.

ascending to erect, 1–3(–4.5) dm, hairs ascending to appressed.

Basal leaves

4–10(–12) × 1–2 cm;

leaflets 3–6(or 7) per side, narrowly obovate, 5–12(–15) × 4–8(–10) mm, 1/2–3/4 as wide as long, divided ± 1/4 to midrib into 5–10 oblong to broadly obovate teeth, sparsely pilose especially on midveins and margins.

± planar, 3–8(–9) × 0.7–1.5 cm;

stipules entire;

leaflets 6–9 per side, separate to ± overlapping at least distally, ± cuneate to obovate, 4–10 × 2–7 mm, 1/2 to nearly as wide as long, divided 1/3 to midrib into 3–6 acute, sometimes obtuse, teeth, densely strigulose, often villous marginally or at least apically.

Cauline leaves

1–4.

(2 or)3–6.

Inflorescences

± congested to open, most flowers arranged in glomerules.

Pedicels

(3–)5–15 mm.

(1–)2–4(–6) mm.

Flowers

15 mm diam.;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm, ± 2/3 length of sepals;

hypanthium 0.6–1 × 2.5–4 mm, less than 1/3 as deep as wide;

sepals ± spreading to reflexed, lanceolate to narrowly elliptic, 3.5–6 mm;

petals elliptic to obovate, 4–7 × 3 mm, apex obtuse to rounded to truncate or slightly emarginate;

filaments 1–3 × 0.6–1.3 mm, anthers 0.6–1 mm;

carpels (17–)20–50;

styles 1.5–2.5 mm.

10 mm diam.;

epicalyx bractlets elliptic-lanceolate, (1.5–)1.8–3 × 0.5–1 mm, ± 2/3 length of sepals, entire;

hypanthium 1–1.5 × 2–4 mm, less than 1/2 as deep as wide, interior slightly pilose;

sepals spreading to ± reflexed, lanceolate, (2.5–)3–4 mm;

petals oblong to oblanceolate, 3–5.5 × 1.5–2 mm, apex rounded to slightly emarginate;

filaments 0.5–2 × 0.3–0.4 mm, anthers (0.4–)0.5–0.8 mm;

carpels 10–20(–27);

styles (1–)1.5–2 mm.

Achenes

grayish, 1.3–1.5 mm, finely reticulate.

brown to dark brown, 1–1.3 mm, usually faintly roughened.

2n

= 28.

Horkelia parryi

Horkelia bolanderi

Phenology Flowering late spring–early summer. Flowering summer.
Habitat Chaparral, pine-oak woodlands, primarily on Ione Formation, rarely on schist or limestone Edges of vernal lakes, meadows, seeps, in grasslands, conifer woodlands
Elevation 80–900 m (300–3000 ft) 400–1100 m (1300–3600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Horkelia parryi is known from the foothills of the western Sierra Nevada in Amador, Calaveras, and El Dorado counties, and is disjunct in Mariposa County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

W. L. Jepson ([1923–1925]) and P. A. Munz and I. M. Johnston (1925) independently adopted an expanded circumscription of Potentilla bolanderi that included four species of Horkelia recognized by P. A. Rydberg (1908c): H. bernardina, H. bolanderi, H. clevelandii, and H. rydbergii. These were treated as three varieties, which D. D. Keck (1938) converted to subspecies as H. bolanderi subsp. bolanderi, subsp. clevelandii, and subsp. parryi, the last encompassing H. bernardina and H. rydbergii. These subspecies were restored to species rank by B. Ertter (1993c) as H. bolanderi, H. clevelandii, and H. rydbergii, along with the establishment of H. yadonii to accommodate populations from Monterey County assigned to subsp. parryi by Keck.

As now circumscribed, Horkelia bolanderi is known from five locations in the northern Coast Ranges in Colusa, Lake, and Mendocino counties; records from more southerly localities would be referenced to one of the segregate species discussed above.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 258. FNA vol. 9, p. 251.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Parryae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Synonyms Potentilla parryi Potentilla bolanderi
Name authority Greene: Bull. Calif. Acad. Sci. 2: 416. (1887) A. Gray: Proc. Amer. Acad. Arts 7: 338. (1868)
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