Horkelia marinensis |
Horkelia parryi |
|
---|---|---|
Point Reyes horkelia |
chamise horkelia, Parry horkelia, Parry's horkelia |
|
Habit | Plants matted, green to grayish. | Plants forming open mats. |
Stems | ± decumbent to ascending, 1–3.5(–4.5) dm, hairs spreading to ascending. |
ascending to erect, 1–3(–3.5) dm. |
Basal leaves | ± planar to nearly cylindric, 4–12(–15) × 1–2.5(–3) cm; stipules entire or basally lobed; leaflets 5–10(–12) per side, ± overlapping especially distally, cuneate, 7–12 × 5–10 mm, nearly as wide as long, divided 1/3–1/2 to midrib into (3–)5–10 acute to obtuse teeth, villous. |
4–10(–12) × 1–2 cm; leaflets 3–6(or 7) per side, narrowly obovate, 5–12(–15) × 4–8(–10) mm, 1/2–3/4 as wide as long, divided ± 1/4 to midrib into 5–10 oblong to broadly obovate teeth, sparsely pilose especially on midveins and margins. |
Cauline leaves | 3–5. |
1–4. |
Inflorescences | usually congested, flowers usually arranged in dense corymbiform clusters. |
|
Pedicels | 1–6 mm. |
(3–)5–15 mm. |
Flowers | 10–15 mm diam.; epicalyx bractlets lanceolate, 2.5–4.5 × 1 mm, ± 2/3 length of sepals, entire; hypanthium 1.2–1.5 × 4–5 mm, less than 1/2 as deep as wide, interior pilose; sepals ± reflexed, lanceolate, 3–6 mm; petals narrowly oblanceolate to narrowly oblong, 4–7 × 2 mm, apex obtuse to rounded, rarely truncate; filaments 1–2.8 × 0.2–0.6 mm, anthers 0.6–0.9 mm; carpels 20–30; styles 2–4 mm. |
15 mm diam.; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm, ± 2/3 length of sepals; hypanthium 0.6–1 × 2.5–4 mm, less than 1/3 as deep as wide; sepals ± spreading to reflexed, lanceolate to narrowly elliptic, 3.5–6 mm; petals elliptic to obovate, 4–7 × 3 mm, apex obtuse to rounded to truncate or slightly emarginate; filaments 1–3 × 0.6–1.3 mm, anthers 0.6–1 mm; carpels (17–)20–50; styles 1.5–2.5 mm. |
Achenes | brown, 1.5–2 mm, smooth to slightly rugose. |
grayish, 1.3–1.5 mm, finely reticulate. |
2n | = 56. |
= 28. |
Horkelia marinensis |
Horkelia parryi |
|
Phenology | Flowering summer. | Flowering late spring–early summer. |
Habitat | Sandy coastal flats, stabilized dunes | Chaparral, pine-oak woodlands, primarily on Ione Formation, rarely on schist or limestone |
Elevation | 0–800 m (0–2600 ft) | 80–900 m (300–3000 ft) |
Distribution |
CA
|
CA
|
Discussion | Of conservation concern. Horkelia marinensis occurs in three population clusters along the coast of California: near Fort Bragg and Rockport in Mendocino County, on Point Reyes Peninsula in Marin County, and from Santa Cruz and (historically) San Mateo counties. The last population cluster has often been confused with H. cuneata var. sericea; plants of it have more deeply and apically toothed, cuneate leaflets and longer, silkier hairs. It is also the basis for the attribution by J. H. Thomas (1961) of H. rydbergii (as H. bolanderi subsp. parryi) to the Santa Cruz Mountains. The three population clusters differ somewhat in vestiture, evident glandularity, and leaflet shape and size; further studies may indicate that varietal status is justified. References to H. marinensis in Sonoma County (M. G. Barbour 1970) are based on specimens of Acaena pinnatifida var. californica; reports from southern Mendocino County (G. L. Smith and C. R. Wheeler 1990–1991) are based on misidentifications of H. californica and H. tenuiloba. Horkelia marinensis is by far the most aromatic member of the genus; the odor, which has been compared to that of a beehive, often can be perceived even before the plants are located, especially on sunny days. Plants do relatively well in cultivation, forming spreading mats with flowers that are strongly attractive to both native bees and honeybees. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Horkelia parryi is known from the foothills of the western Sierra Nevada in Amador, Calaveras, and El Dorado counties, and is disjunct in Mariposa County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 253. | FNA vol. 9, p. 258. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Parryae |
Sibling taxa | ||
Synonyms | H. bolanderi var. marinensis, Potentilla kelloggii var. marinensis, P. marinensis | Potentilla parryi |
Name authority | (Elmer) Crum: Lloydia 1: 91. (1939) | Greene: Bull. Calif. Acad. Sci. 2: 416. (1887) |
Web links |