Horkelia hispidula |
Horkelia tridentata |
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White Mountain horkelia, White Mountains. horkelia |
cluster horkelia, three-tooth horkelia |
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Habit | Plants 0.7–3 dm diam. | Plants mostly rosette-forming, grayish. | ||||
Stems | ascending to erect, 1–2.5 dm. |
decumbent to nearly erect, 0.5–4(–7) dm, hairs 1–3 mm proximally, glands absent or sparse distally. |
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Basal leaves | 3–10 × 0.4–0.8 cm; leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid. |
planar, 3–12 × 1–3(–4) cm, usually sericeous to villous, sometimes glabrate adaxially; stipules entire or rarely forked or pectinate; leaflets 2–5(or 6) per side, separate to slightly overlapping, elliptic to cuneate-oblong, (3–)5–20(–30) × 1.5–7(–10) mm, (1/4–)1/3–1/2 as wide as long, divided 1/10–1/6 or less to midrib into (0–)3(or 4) triangular teeth 1 mm wide, these restricted to apex. |
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Cauline leaves | 3–7. |
1–5; stipules 3–12 mm, entire or deeply 3–5-lobed, lanceolate to ovate. |
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Inflorescences | open to congested, flowers arranged in ± capitate glomerules or in ± corymbiform clusters. |
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Pedicels | 2–8(–12) mm. |
(1–)2–6(–10) mm. |
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Flowers | 3–15, 10 mm diam.; epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals; hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose; sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm; petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate; filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm; carpels (10–)12–18(–20); styles 1.8–2.2 mm. |
4–10 mm diam.; epicalyx bractlets linear to narrowly elliptic, 0.8–2(–3) × 0.2–0.4 mm, 1/2–2/3 length of sepals; hypanthium 1–2.5 × 2–5 mm, 1/2 to nearly as deep as wide, interior glabrous or pilose; sepals moderately reflexed, abaxially green to reddish, 1.5–3 mm; petals white, sometimes tinged with pink, linear to broadly oblanceolate, (1–)1.5–4(–4.5) × 0.3–1.5 mm, apex acute to rounded or mucronate; filaments 0.5–2 × 0.1–0.4(–0.8) mm, anthers 0.2–0.5 mm; carpels 5–15; styles 1–2.5 mm. |
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Achenes | brown to dark brown, 1.5–2 mm. |
light brown, 1.5–2.5 mm, rugose. |
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Horkelia hispidula |
Horkelia tridentata |
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Phenology | Flowering summer. | |||||
Habitat | Dry, rocky alpine flats, in subalpine conifer woodlands | |||||
Elevation | 3000–3400 m (9800–11200 ft) | |||||
Distribution |
CA; NV |
CA; OR
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Discussion | Of conservation concern. When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Rydberg established Horkelia flavescens on the basis of its more congested inflorescence, purportedly yellowish petals, and broader filaments. As noted by D. D. Keck (1938), the petals are usually white, at least when fresh, and the other characters are highly variable; he reduced H. flavescens to a subspecies of H. tridentata, which is here treated at the rank of variety. In general, var. flavescens tends to be more decumbent with a single, loosely congested flower cluster and broader petals; these characters completely overlap and can vary independently. The presence or absence of hairs on the hypanthium interior is the single most reliably diagnostic character and is the primary determiner of the geographic distributions presented here. The exception is a cluster of populations from the extreme southern end of the coastal range of var. flavescens in Colusa, Lake, and Mendocino counties, California, which have glabrous hypanthia but otherwise have all the diagnostic characters of var. flavescens. Although provisionally included here within var. flavescens, these populations may be better treated as a separate variety. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 249. | FNA vol. 9, p. 269. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Potentilla hispidula | |||||
Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908) | Torrey: in War Department [U.S.], Pacif. Railr. Rep. 4(5): 84, plate 6. (1857) | ||||
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