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White Mountain horkelia, White Mountains. horkelia

carrot leafed horkelia, carrot-leaf horkelia

Habit Plants 0.7–3 dm diam. Plants rosette-forming to tufted, rarely ± matted, grayish or green.
Stems

ascending to erect, 1–2.5 dm.

ascending to erect, 1.5–3.5 dm, hairs 2–3 mm proximally, glands sparse to dense distally.

Basal leaves

3–10 × 0.4–0.8 cm;

leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid.

weakly planar to loosely cylindric, (4–)5–12(–17) × 0.8–3(–4) cm, usually sericeous to villous, sometimes glabrate;

stipules pinnately divided into 3–7 linear to filiform lobes;

leaflets 5–10 per side, ± overlapping, obovate to broadly obcordate, 5–15(–25) × 5–20 mm, ± as wide as long, divided 3/4+ to midrib into (0–)2–15 linear to oblanceolate lobes 0.4–2(–3) mm wide, these not restricted to apex.

Cauline leaves

3–7.

3–6;

stipules 8–15 mm, deeply 3–7-lobed proximally, 1–3-lobed distally.

Inflorescences

open to ± congested, flowers arranged individually or in ± corymbiform clusters.

Pedicels

2–8(–12) mm.

2–10(–20) mm.

Flowers

3–15, 10 mm diam.;

epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals;

hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose;

sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm;

petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate;

filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm;

carpels (10–)12–18(–20);

styles 1.8–2.2 mm.

10–15 mm diam.;

epicalyx bractlets linear to linear-lanceolate, 2.5–3.5 × 0.5 mm, 2/3 to ± equal to length of sepals;

hypanthium 1–1.5 × 3.5–5 mm, less than 1/2 as deep as wide, interior glabrous;

sepals ± spreading, abaxially green, 3.5–7.5 mm;

petals white to cream, often drying yellowish, cuneate to obcordate, (3.5–)4–8 × 2–8 mm, apex ± emarginate, sometimes rounded;

filaments 1–3 × 0.5–1.5 mm, anthers 0.5–0.8 mm;

carpels 5–15, styles 2–4 mm.

Achenes

brown to dark brown, 1.5–2 mm.

dark brown, 2.4–3 mm, smooth to ± roughened.

Horkelia hispidula

Horkelia daucifolia

Phenology Flowering summer.
Habitat Dry, rocky alpine flats, in subalpine conifer woodlands
Elevation 3000–3400 m (9800–11200 ft)
Distribution
from FNA
CA; NV
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

D. D. Keck (1938) decided that the leaflet characters previously used to recognize segregates of Horkelia daucifolia were random variables that have not become localized into geographic subunits. At the same time, he established subsp. latior D. D. Keck to accommodate a single collection with leaves approaching those of H. sericata, with the speculation that the population was an amphiploid derivative of the two species. Subsequent collections from the same general area (Scott Mountain, Siskiyou County, California) provide a full range of variation between the extreme with exceptionally wide leaflet lobes and the typical form occurring at the base of the mountain. This variant of the species is accordingly not recognized here, but the extremes with narrower, more numerous leaflet lobes are recognized as varieties that coincide with major river drainages (B. Ertter and J. L. Reveal 2007).

In the descriptions below, the pedicels of all varieties are sparsely pilose in addition to being puberulent or not.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet lobes 2–5(–8), linear to oblanceolate, 1–2(–3) mm wide; petals cuneate to obovate, 2–4 mm wide.
var. daucifolia
1. Leaflet lobes (2–)5–15, linear, 0.4–1 mm wide; petals broadly obovate to obcordate, 4–8 mm wide
→ 2
2. Pedicels 2–7(–10) mm, puberulent, eglandular; Oregon.
var. caruifolia
2. Pedicels 5–10(–20) mm, not puberulent, minutely glandular; California.
var. indicta
Source FNA vol. 9, p. 249. FNA vol. 9, p. 266.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Hispidulae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Subordinate taxa
H. daucifolia var. caruifolia, H. daucifolia var. daucifolia, H. daucifolia var. indicta
Synonyms Potentilla hispidula Potentilla daucifolia
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908) (Greene) Rydberg: Bull. Torrey Bot. Club 25: 55. (1898)
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