Horkelia hispidula |
Horkelia cuneata |
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White Mountain horkelia, White Mountains. horkelia |
wedge leaf horkelia |
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Habit | Plants 0.7–3 dm diam. | Plants matted or tufted, green to grayish. | ||||||||
Stems | ascending to erect, 1–2.5 dm. |
decumbent to erect, (1–)2–6(–7.5) dm, hairs ascending to appressed, sometimes spreading. |
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Basal leaves | 3–10 × 0.4–0.8 cm; leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid. |
planar, (5–)10–30 × 1.5–4(–5) cm; stipules entire or basally lobed; leaflets 5–12 per side, separate to slightly overlapping, ± elliptic to obovate or oblanceolate, 5–25(–30) × 5–15(–25) mm, ± 1/2 to nearly as wide as long, divided less than 1/3 to midrib into (5–)10–15 oblanceolate to obovate teeth, sparsely pilose to glabrate or sericeous. |
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Cauline leaves | 3–7. |
1–3(–5). |
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Inflorescences | ± open to congested, flowers arranged individually and/or in glomerules. |
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Pedicels | 2–8(–12) mm. |
1–30(–40) mm. |
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Flowers | 3–15, 10 mm diam.; epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals; hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose; sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm; petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate; filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm; carpels (10–)12–18(–20); styles 1.8–2.2 mm. |
12–17 mm diam.; epicalyx bractlets lanceolate to ovate, 3–5 × 1.5–3 mm, 2/3–3/4 length of sepals, entire; hypanthium 1.5–2 × 4–7 mm, less than 1/2 as deep as wide, interior glabrous or pilose in a narrow ring; sepals ± reflexed to spreading, lanceolate, 4–6.5 mm; petals oblanceolate to narrowly obovate, 4–8 × 1.5–4 mm, apex obtuse to rounded; filaments 1–3 × 0.5–2 mm, anthers 0.7–1.1 mm; carpels (30–)40–60(–80); styles 2–3 mm. |
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Achenes | brown to dark brown, 1.5–2 mm. |
brown, 1.5–1.8 mm, smooth or minutely rugose. |
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Horkelia hispidula |
Horkelia cuneata |
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Phenology | Flowering summer. | |||||||||
Habitat | Dry, rocky alpine flats, in subalpine conifer woodlands | |||||||||
Elevation | 3000–3400 m (9800–11200 ft) | |||||||||
Distribution |
CA; NV |
CA
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Discussion | Of conservation concern. When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). As noted by D. D. Keck (1938) and B. Ertter (1997b), recognition of subunits within Horkelia cuneata is complicated by the abundance of intermediates, to the extent that a complete transition exists between the decumbent, densely hairy coastal extreme with a congested inflorescence, represented by var. sericea, and the erect, green-glandular interior extreme with an open inflorescence, represented by var. puberula. The traditional division into three infraspecific units is maintained here to emphasize the distinctiveness of the extremes, which would qualify as specifically distinct were they not ends of a continuum. The extremes are furthermore threatened by ongoing decimation of suitable habitats at the northern (San Francisco Bay Area) and southern (greater Los Angeles) ends of the species range. Determining the correct name of Horkelia cuneata if treated as a species of Potentilla is complicated. Potentilla cuneata (Lindley) Baillon ex Munz & I. M. Johnston (1925) and P. puberula Greene are both later homonyms, while P. multijuga Lehmann has been conserved with a conserved type to maintain established usage for an unrelated species of Potentilla (B. Ertter and J. L. Reveal 2008). J. T. Howell (1945) adopted P. lindleyi; however, as noted by R. F. Hoover (1966), P. kelloggii has priority at species rank by a matter of months. The latter is accordingly the correct name in Potentilla for H. cuneata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 249. | FNA vol. 9, p. 254. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Potentilla hispidula | Potentilla kelloggii var. cuneata, P. lindleyi | ||||||||
Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908) | Lindley: Edwards’s Bot. Reg. 23: sub plate 1997. (1837) | ||||||||
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