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White Mountain horkelia, White Mountains. horkelia

Bolander's horkelia, border horkelia

Habit Plants 0.7–3 dm diam. Plants openly matted, grayish.
Stems

ascending to erect, 1–2.5 dm.

ascending to erect, 1–3(–4.5) dm, hairs ascending to appressed.

Basal leaves

3–10 × 0.4–0.8 cm;

leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid.

± planar, 3–8(–9) × 0.7–1.5 cm;

stipules entire;

leaflets 6–9 per side, separate to ± overlapping at least distally, ± cuneate to obovate, 4–10 × 2–7 mm, 1/2 to nearly as wide as long, divided 1/3 to midrib into 3–6 acute, sometimes obtuse, teeth, densely strigulose, often villous marginally or at least apically.

Cauline leaves

3–7.

(2 or)3–6.

Inflorescences

± congested to open, most flowers arranged in glomerules.

Pedicels

2–8(–12) mm.

(1–)2–4(–6) mm.

Flowers

3–15, 10 mm diam.;

epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals;

hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose;

sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm;

petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate;

filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm;

carpels (10–)12–18(–20);

styles 1.8–2.2 mm.

10 mm diam.;

epicalyx bractlets elliptic-lanceolate, (1.5–)1.8–3 × 0.5–1 mm, ± 2/3 length of sepals, entire;

hypanthium 1–1.5 × 2–4 mm, less than 1/2 as deep as wide, interior slightly pilose;

sepals spreading to ± reflexed, lanceolate, (2.5–)3–4 mm;

petals oblong to oblanceolate, 3–5.5 × 1.5–2 mm, apex rounded to slightly emarginate;

filaments 0.5–2 × 0.3–0.4 mm, anthers (0.4–)0.5–0.8 mm;

carpels 10–20(–27);

styles (1–)1.5–2 mm.

Achenes

brown to dark brown, 1.5–2 mm.

brown to dark brown, 1–1.3 mm, usually faintly roughened.

Horkelia hispidula

Horkelia bolanderi

Phenology Flowering summer. Flowering summer.
Habitat Dry, rocky alpine flats, in subalpine conifer woodlands Edges of vernal lakes, meadows, seeps, in grasslands, conifer woodlands
Elevation 3000–3400 m (9800–11200 ft) 400–1100 m (1300–3600 ft)
Distribution
from FNA
CA; NV
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

W. L. Jepson ([1923–1925]) and P. A. Munz and I. M. Johnston (1925) independently adopted an expanded circumscription of Potentilla bolanderi that included four species of Horkelia recognized by P. A. Rydberg (1908c): H. bernardina, H. bolanderi, H. clevelandii, and H. rydbergii. These were treated as three varieties, which D. D. Keck (1938) converted to subspecies as H. bolanderi subsp. bolanderi, subsp. clevelandii, and subsp. parryi, the last encompassing H. bernardina and H. rydbergii. These subspecies were restored to species rank by B. Ertter (1993c) as H. bolanderi, H. clevelandii, and H. rydbergii, along with the establishment of H. yadonii to accommodate populations from Monterey County assigned to subsp. parryi by Keck.

As now circumscribed, Horkelia bolanderi is known from five locations in the northern Coast Ranges in Colusa, Lake, and Mendocino counties; records from more southerly localities would be referenced to one of the segregate species discussed above.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 249. FNA vol. 9, p. 251.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Hispidulae Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia
Sibling taxa
H. bolanderi, H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
H. californica, H. clevelandii, H. congesta, H. cuneata, H. daucifolia, H. fusca, H. hendersonii, H. hispidula, H. howellii, H. marinensis, H. parryi, H. rydbergii, H. sericata, H. tenuiloba, H. tridentata, H. truncata, H. tularensis, H. wilderae, H. yadonii
Synonyms Potentilla hispidula Potentilla bolanderi
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908) A. Gray: Proc. Amer. Acad. Arts 7: 338. (1868)
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