Horkelia fusca |
Horkelia hispidula |
|||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
dusky, dusky horkelia, horkelia, pinewoods, pinewoods horkelia, pink pinwheels, tawny horkelia |
White Mountain horkelia, White Mountains. horkelia |
|||||||||||||||||||||||||
Habit | Plants 0.7–3 dm diam. | |||||||||||||||||||||||||
Stems | ascending to erect, 1–2.5 dm. |
|||||||||||||||||||||||||
Basal leaves | (3–)4–20(–40) × (1–)1.5–4(–7) cm; leaflets narrowly cuneate to obovate to flabellate, 5–30(–35) × 2–20(–30) mm, 1/3 as wide to wider than long, divided into linear or oblanceolate to obovate teeth or lobes, sparsely to ± densely short-villous or hirsute, sometimes glabrate. |
3–10 × 0.4–0.8 cm; leaflets (6–)10–14 per side, ± overlapping at least distally, cuneate to flabellate, 2.5–4(–6) mm, divided 3/4+ to midrib into 3–6 oblanceolate to obovate lobes, hispid. |
||||||||||||||||||||||||
Cauline leaves | 1–5(or 6). |
3–7. |
||||||||||||||||||||||||
Pedicels | 2–8(–12) mm. |
|||||||||||||||||||||||||
Flowers | 5–12 mm diam.; epicalyx bractlets 1–3 mm, 1/2 length of to nearly equal to sepals; hypanthium 1–3 × 2–4 mm, 1/2 to nearly as deep as wide; sepals spreading to ± reflexed, narrowly to broadly lanceolate, (1.7–)2–4(–4.5) mm; petals 2–6(–6.5) mm; filaments 0.2–1.5 × (0.2–)0.4–0.6(–1) mm, anthers 0.4–0.6 mm; styles 0.9–1.5 mm. |
3–15, 10 mm diam.; epicalyx bractlets linear to lanceolate, 1.5–3 × 0.2–0.5 mm, ± 2/3 length of sepals; hypanthium 1.8–3 × 3–4 mm, ± 1/2 as deep as wide, interior sparsely pilose; sepals reflexed, broadly lanceolate, 2.5–4(–5) mm, hairs stiff, 0.5 mm; petals not pink-tinged, oblanceolate to oblong or narrowly elliptic, 2.5–5 mm, apex rounded to truncate, sometimes slightly emarginate or mucronate; filaments white, 0.5–2 × 0.4–0.6 mm, glabrous or sparsely pilose adaxially, anthers 0.5–0.9 mm; carpels (10–)12–18(–20); styles 1.8–2.2 mm. |
||||||||||||||||||||||||
Achenes | brown. |
brown to dark brown, 1.5–2 mm. |
||||||||||||||||||||||||
Horkelia fusca |
Horkelia hispidula |
|||||||||||||||||||||||||
Phenology | Flowering summer. | |||||||||||||||||||||||||
Habitat | Dry, rocky alpine flats, in subalpine conifer woodlands | |||||||||||||||||||||||||
Elevation | 3000–3400 m (9800–11200 ft) | |||||||||||||||||||||||||
Distribution |
CA; ID; NV; OR; WA; WY
|
CA; NV |
||||||||||||||||||||||||
Discussion | Varieties 7 (7 in the flora). Horkelia fusca represents the primary radiation of the genus beyond the California Floristic Province. Within California, H. fusca occurs in the Sierra Nevada and mountains of northern California, and it is the only representative of the genus extending beyond California and Oregon into Washington, Idaho, Nevada [with the possible exception of H. tridentata (10e. sect. Tridentatae) in Washoe County], and, questionably, Wyoming. As here circumscribed, Horkelia fusca is the most diverse, most widely distributed species in the genus, with significant additional work needed to fully elucidate its variation patterns. The treatment presented here differs from that of D. D. Keck (1938) and B. Ertter (1993d) in using the rank variety instead of subspecies, circumscribing var. capitata more narrowly, and transferring the application of var. pseudocapitata from what is here called var. brownii to the bulk of what had been subsp. capitata (Lindley) D. D. Keck (B. Ertter and J. L. Reveal 2007). There are two types of basal leaves in plants of Horkelia fusca. The ephemeral early-season leaves have leaflets that tend to be broadly cuneate-obovate, shallowly toothed, densely glandular but otherwise sparsely hairy, and deeply veined. The leaf features described below are drawn from the more persistent, mid season leaves that predominate at peak flowering and differ more strongly among varieties. Petals of first-formed flowers are often larger than average; end-of-season petals can be smaller than average. Although Montana is sometimes included in the range of Horkelia fusca, such references are based only on potential occurrence (W. E. Booth and J. C. Wright 1959). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. When describing Horkelia hispidula, Rydberg associated it with H. sericata in his group Sericatae. The species is known only from the White Mountains of California and adjacent Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||
Source | FNA vol. 9, p. 259. | FNA vol. 9, p. 249. | ||||||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Hispidulae | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Potentilla douglasii | Potentilla hispidula | ||||||||||||||||||||||||
Name authority | Lindley: Edwards’s Bot. Reg. 23: plate 1997. (1837) | Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 278. (1908) | ||||||||||||||||||||||||
Web links |