Horkelia fusca
Horkelia daucifolia
dusky, horkelia, pinewoods, pinewoods horkelia, tawny horkelia
carrot leafed horkelia, carrot-leaf horkelia
ascending to erect, 1.5–3.5 dm, hairs 2–3 mm proximally, glands sparse to dense distally.
(3–)4–20(–40) × (1–)1.5–4(–7) cm;
leaflets narrowly cuneate to obovate to flabellate, 5–30(–35) × 2–20(–30) mm, 1/3 as wide to wider than long, divided into linear or oblanceolate to obovate teeth or lobes, sparsely to ± densely short-villous or hirsute, sometimes glabrate.
weakly planar to loosely cylindric, (4–)5–12(–17) × 0.8–3(–4) cm, usually sericeous to villous, sometimes glabrate;
stipules pinnately divided into 3–7 linear to filiform lobes;
leaflets 5–10 per side, ± overlapping, obovate to broadly obcordate, 5–15(–25) × 5–20 mm, ± as wide as long, divided 3/4+ to midrib into (0–)2–15 linear to oblanceolate lobes 0.4–2(–3) mm wide, these not restricted to apex.
1–5(or 6).
3–6;
stipules 8–15 mm, deeply 3–7-lobed proximally, 1–3-lobed distally.
open to ± congested, flowers arranged individually or in ± corymbiform clusters.
2–10(–20) mm.
5–12 mm diam.;
epicalyx bractlets 1–3 mm, 1/2 length of to nearly equal to sepals;
hypanthium 1–3 × 2–4 mm, 1/2 to nearly as deep as wide;
sepals spreading to ± reflexed, narrowly to broadly lanceolate, (1.7–)2–4(–4.5) mm;
petals 2–6(–6.5) mm;
filaments 0.2–1.5 × (0.2–)0.4–0.6(–1) mm, anthers 0.4–0.6 mm;
styles 0.9–1.5 mm.
10–15 mm diam.;
epicalyx bractlets linear to linear-lanceolate, 2.5–3.5 × 0.5 mm, 2/3 to ± equal to length of sepals;
hypanthium 1–1.5 × 3.5–5 mm, less than 1/2 as deep as wide, interior glabrous;
sepals ± spreading, abaxially green, 3.5–7.5 mm;
petals white to cream, often drying yellowish, cuneate to obcordate, (3.5–)4–8 × 2–8 mm, apex ± emarginate, sometimes rounded;
filaments 1–3 × 0.5–1.5 mm, anthers 0.5–0.8 mm;
carpels 5–15, styles 2–4 mm.
brown.
dark brown, 2.4–3 mm, smooth to ± roughened.
Horkelia fusca
Horkelia daucifolia
Varieties 7 (7 in the flora).
Horkelia fusca represents the primary radiation of the genus beyond the California Floristic Province. Within California, H. fusca occurs in the Sierra Nevada and mountains of northern California, and it is the only representative of the genus extending beyond California and Oregon into Washington, Idaho, Nevada [with the possible exception of H. tridentata (10e. sect. Tridentatae) in Washoe County], and, questionably, Wyoming.
As here circumscribed, Horkelia fusca is the most diverse, most widely distributed species in the genus, with significant additional work needed to fully elucidate its variation patterns. The treatment presented here differs from that of D. D. Keck (1938) and B. Ertter (1993d) in using the rank variety instead of subspecies, circumscribing var. capitata more narrowly, and transferring the application of var. pseudocapitata from what is here called var. brownii to the bulk of what had been subsp. capitata (Lindley) D. D. Keck (B. Ertter and J. L. Reveal 2007).
There are two types of basal leaves in plants of Horkelia fusca. The ephemeral early-season leaves have leaflets that tend to be broadly cuneate-obovate, shallowly toothed, densely glandular but otherwise sparsely hairy, and deeply veined. The leaf features described below are drawn from the more persistent, mid season leaves that predominate at peak flowering and differ more strongly among varieties.
Petals of first-formed flowers are often larger than average; end-of-season petals can be smaller than average.
Although Montana is sometimes included in the range of Horkelia fusca, such references are based only on potential occurrence (W. E. Booth and J. C. Wright 1959).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 3 (3 in the flora).
D. D. Keck (1938) decided that the leaflet characters previously used to recognize segregates of Horkelia daucifolia were random variables that have not become localized into geographic subunits. At the same time, he established subsp. latior D. D. Keck to accommodate a single collection with leaves approaching those of H. sericata, with the speculation that the population was an amphiploid derivative of the two species. Subsequent collections from the same general area (Scott Mountain, Siskiyou County, California) provide a full range of variation between the extreme with exceptionally wide leaflet lobes and the typical form occurring at the base of the mountain. This variant of the species is accordingly not recognized here, but the extremes with narrower, more numerous leaflet lobes are recognized as varieties that coincide with major river drainages (B. Ertter and J. L. Reveal 2007).
In the descriptions below, the pedicels of all varieties are sparsely pilose in addition to being puberulent or not.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflets divided ± 1/2–3/4+ to midrib | → 2 |
1. Leaflets divided less than 1/2 to midrib | → 4 |
2. Inflorescences usually with flowers arranged individually; sw Oregon. | var. filicoides |
2. Inflorescences composed of 3–20-flowered glomerules, (flowers rarely arranged individually in var. fusca); ne California, nw Oregon, sw Washington | → 3 |
3. Leaflets divided ± 1/2 to midrib, 5–18(–20) × 4–15 mm; petals 3–5 mm; filaments 0.5–1.3 mm, longer than wide; nw Oregon, sw Washington. | var. fusca |
3. Leaflets divided 3/4+ to midrib, 5–10(–15) × 2–10(–20) mm; petals 2–3(–4) mm; filaments 0.2–0.5 mm, wider than long; ne California. | var. tenella |
4. Leaves grayish, surfaces often obscured, ± densely short-villous; leaflets narrowly to broadly cuneate, usually 1/3–1/2 as wide as long; ne California, adjacent Oregon and Nevada. | var. brownii |
4. Leaves green, surfaces not obscured, ± short-hirsute, villous, or glabrate; leaflets cuneate to broadly obovate, 1/2 to wider than long; California to Washington, Idaho, Wyoming | → 5 |
5. Inflorescences dark reddish purple, densely congested, bracts acuminate-lobed, obscuring pedicels and flowers even at maturity, glandular hairs usually conspicuously red-septate; epicalyx bractlets (2–)3.5–5(–6) mm; cauline leaves 3–6, leaflets of proximalmost 3–5 per side; ne Oregon, se Washington, n Idaho. | var. capitata |
5. Inflorescences green to reddish purple, congested to open, bracts acute-lobed, not obscuring pedicels and flowers at maturity, glandular hairs not or obscurely red-septate; epicalyx bractlets 1–3.5(–4) mm; cauline leaves 1–4(or 5), leaflets of proximalmost 2–4(or 5) per side; Cascade-Sierra axis of Oregon, California, e to Wyoming and n Nevada | → 6 |
6. Petals (3–)4–6 mm; leaflets 10–30(–35) × (5–)10–20(–25) mm; epicalyx bractlets 2–3.5(–4) mm; mountains in and bordering n Intermountain Region, ne California, n Nevada, se Oregon, s Idaho. | var. pseudocapitata |
6. Petals 2–4.5(–6.5) mm; leaflets 5–15(–20) × (2–)5–10(–15) mm; epicalyx bractlets 1–2(–3) mm; Sierra Nevada, California and Nevada, to Cascade Ranges, c Oregon, and mountains of c Idaho, adjacent Wyoming. | var. parviflora |
1. Leaflet lobes 2–5(–8), linear to oblanceolate, 1–2(–3) mm wide; petals cuneate to obovate, 2–4 mm wide. | var. daucifolia |
1. Leaflet lobes (2–)5–15, linear, 0.4–1 mm wide; petals broadly obovate to obcordate, 4–8 mm wide | → 2 |
2. Pedicels 2–7(–10) mm, puberulent, eglandular; Oregon. | var. caruifolia |
2. Pedicels 5–10(–20) mm, not puberulent, minutely glandular; California. | var. indicta |
CA
OR
WA
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