Horkelia fusca var. pseudocapitata
Horkelia sect. Capitatae
intermountain horkelia, tawny horkelia
3–6 dm.
ascending to erect, (0.6–)1–6(–9) dm.
green, 10–35(–40) cm;
leaflets 4–6(or 7) per side, narrowly to broadly obovate, 10–30(–35) × (5–)10–20(–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate.
planar;
stipules entire;
leaflets 3–8(–15) per side, separate, sometimes ± overlapping, divided 1/5–3/4+ to midrib into (3–)5–15 teeth or lobes not restricted to apex.
1–4(or 5);
leaflets of proximalmost 2 or 3(–5) per side.
green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular hairs not or obscurely red-septate;
bracts acute-lobed, not obscuring pedicels and flowers at maturity.
open to ± congested, flowers usually arranged in ± capitate glomerules, arranged individually in H. fusca var. filicoides.
remaining ± straight, outermost sometimes ± reflexed in congested inflorescences, 1–3(–10) mm.
epicalyx bractlets 2–3.5(–4) mm;
hypanthium 2 × 2.5–4 mm;
petals (3–)4–6 mm;
filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm;
styles 1–1.5 mm.
epicalyx bractlets linear, 0.2–0.3(–0.5) mm wide, entire;
hypanthium interior glabrous;
sepals acute to acuminate;
petals white to pale pink, often veined with pink to rose, ± oblanceolate to cuneate, apex emarginate to truncate or rounded;
filaments white, glabrous, anthers longer than wide;
carpels 10–25.
1.6–1.8 mm.
1–1.8 mm, smooth.
= 28.
Horkelia fusca var. pseudocapitata
Horkelia sect. Capitatae
The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007).
As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 1.
Rydberg recognized seven species in his group Capitatae, treated here as intergrading variation within a single species. Plants of the section share the characteristic Horkelia odor, glandularity, and planar leaves of sect. Horkelia, but differ in the combination of relatively small, short-pedicelled flowers that (except for var. filicoides) are most commonly aggregated into one or more capitate, purple-suffused glomerules, with linear epicalyx bractlets, oblanceolate-cuneate petals that are often pink-tinged, and relatively short, broad filaments.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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