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Photo is of parent taxon

intermountain horkelia, tawny horkelia

Photo is of parent taxon

delicate horkelia, pinewoods horkelia, tawny horkelia

Stems

3–6 dm.

(0.6–)1–4(–5) dm.

Basal leaves

green, 10–35(–40) cm;

leaflets 4–6(or 7) per side, narrowly to broadly obovate, 10–30(–35) × (5–)10–20(–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate.

green, (3–)4–18(–27) cm;

leaflets 8–15 per side, cuneate to flabellate, 5–10(–15) × 2–10(–20) mm, 3/4 as wide to wider than long, divided 3/4+ to midrib into 5–15 lobes or teeth, surfaces not obscured, sparsely short-hirsute to glabrate.

Cauline leaves

1–4(or 5);

leaflets of proximalmost 2 or 3(–5) per side.

2–4(or 5);

leaflets of proximalmost 3–6 per side.

Inflorescences

green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular hairs not or obscurely red-septate;

bracts acute-lobed, not obscuring pedicels and flowers at maturity.

green to ± reddish, congested to open, comprising no more than 1/4 of stem, composed of 3–10(–20)-flowered glomerules, glandular hairs sometimes red-septate;

bracts acute- to acuminate-lobed, sometimes partly obscuring pedicels and flowers at maturity.

Flowers

epicalyx bractlets 2–3.5(–4) mm;

hypanthium 2 × 2.5–4 mm;

petals (3–)4–6 mm;

filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm;

styles 1–1.5 mm.

epicalyx bractlets 1.5–2 mm;

hypanthium 1.5–2 × 2–3 mm;

petals 2–3(–4) mm;

filaments 0.2–0.5 mm, wider than long, anthers 0.5 mm;

styles 1 mm.

Achenes

1.6–1.8 mm.

1.2–1.5 mm.

2n

= 28.

= 28.

Horkelia fusca var. pseudocapitata

Horkelia fusca var. tenella

Phenology Flowering summer. Flowering summer.
Habitat Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil Dry meadow edges, open conifer woodlands, mainly on volcanic soil
Elevation 900–2300 m (3000–7500 ft) 1200–2200 m (3900–7200 ft)
Distribution
from FNA
CA; ID; NV; OR
from FNA
CA
[BONAP county map]
Discussion

The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007).

As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of var. tenella have the most deeply and consistently divided leaflets, and the variety is the only one with such leaflets that occurs in California. It is most common in open lodgepole pine forests at the southern end of the Cascade Range in northeastern California. Populations outside this core area tend to intergrade with var. brownii or var. parviflora. A collection from west-central Tehama County (Bracelin 405, UC) matches this variety, but from an atypical habitat well outside the core range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 262. FNA vol. 9, p. 261.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca
Sibling taxa
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. filicoides, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. tenella
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. filicoides, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. pseudocapitata
Synonyms H. pseudocapitata, H. fusca subsp. pseudocapitata, Potentilla douglasii var. pseudocapitata H. fusca subsp. tenella, Potentilla douglasii var. tenella
Name authority (Rydberg ex Howell) M. Peck: Man. Pl. Oregon, 398. (1941) S. Watson: in W. H. Brewer et al., Bot. California 1: 181. (1876)
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