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Photo is of parent taxon

intermountain horkelia, tawny horkelia

Photo is of parent taxon

small-flower horkelia, tawny horkelia

Stems

3–6 dm.

1–4.5 dm.

Basal leaves

green, 10–35(–40) cm;

leaflets 4–6(or 7) per side, narrowly to broadly obovate, 10–30(–35) × (5–)10–20(–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate.

green, 4–15(–18) cm;

leaflets 4–8 per side, cuneate to broadly obovate, 5–15(–20) × (2–)5–10(–15) mm, 1/2 as wide to wider than long, divided 1/4–1/2 to midrib into 4–6(–10) teeth, surfaces not obscured, sparsely to moderately hirsute or villous, sometimes glabrate.

Cauline leaves

1–4(or 5);

leaflets of proximalmost 2 or 3(–5) per side.

1–3(or 4);

leaflets of proximalmost 2–4(or 5) per side.

Inflorescences

green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular hairs not or obscurely red-septate;

bracts acute-lobed, not obscuring pedicels and flowers at maturity.

green to purplish, congested to open, usually comprising less than 1/4 of stem, composed of 5–20-flowered glomerules, glandular hairs not red-septate;

bracts acute-lobed, not obscuring pedicels and flowers at maturity.

Flowers

epicalyx bractlets 2–3.5(–4) mm;

hypanthium 2 × 2.5–4 mm;

petals (3–)4–6 mm;

filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm;

styles 1–1.5 mm.

epicalyx bractlets 1–2(–3) mm;

hypanthium 1.5–2 × 2–3.5 mm;

petals 2–4.5(–6.5) mm;

filaments 0.2–1 mm, usually longer than wide, anthers 0.3–0.5 mm;

styles 0.9–1.1 mm.

Achenes

1.6–1.8 mm.

1–1.2 mm.

2n

= 28.

= 28.

Horkelia fusca var. pseudocapitata

Horkelia fusca var. parviflora

Phenology Flowering summer. Flowering summer.
Habitat Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil Dry meadow edges, in conifer woodlands, mainly on volcanic or granitic soil
Elevation 900–2300 m (3000–7500 ft) 800–3300 m (2600–10800 ft)
Distribution
from FNA
CA; ID; NV; OR
from FNA
CA; ID; NV; OR; WY
[BONAP county map]
Discussion

The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007).

As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variety parviflora is the most widespread and polymorphic in the species and might represent the repository for variation that is not accommodated by the other, more tightly circumscribed varieties. Population clusters occur in three discrete areas: the Sierra Nevada of eastern California and adjacent Nevada, the mountains of northern California extending north in the Cascade Range to central Oregon, and the mountains of central Idaho. Each of these three areas has plants that differ slightly from those of the other two.

The most distinct phase is found in the Cascade Range of Oregon, with larger-than-average petals and highly branched inflorescences. Plants from Lost Prairie in eastern Linn County are particularly anomalous, with petals to 6.5 mm; further studies may indicate that these deserve separate recognition.

Plants from the Sierra Nevada tend to have smaller petals, and those in the higher southern Sierra Nevada tend to have capitate or subcapitate inflorescences. From about Lake Tahoe northward, inflorescences tend to be somewhat more open but still congested compared to the Oregon phase.

The Idaho phase, represented by the type of var. parviflora, has petals that run the full range from 2–4 mm, with most individuals having a slightly branched inflorescence. Unlike those of the other two phases, both the hypanthia and sepals are often purple; the hypanthium is otherwise typically greenish with only the sepals themselves a dark purple. The sole Wyoming record (D. R. Goddard 1027, F, UC) has yet to be confirmed by more recent collections in Yellowstone National Park. These plants have pale inflorescences, relatively blunt sepals, and short epicalyx bractlets, differing in these features from the closest populations in central Idaho. For reports from Montana, see species discussion.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 262. FNA vol. 9.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca
Sibling taxa
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. filicoides, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. tenella
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. filicoides, H. fusca var. fusca, H. fusca var. pseudocapitata, H. fusca var. tenella
Synonyms H. pseudocapitata, H. fusca subsp. pseudocapitata, Potentilla douglasii var. pseudocapitata H. parviflora, H. fusca subsp. parviflora, Potentilla andersonii, P. douglasii var. parviflora
Name authority (Rydberg ex Howell) M. Peck: Man. Pl. Oregon, 398. (1941) (Nuttall ex Hooker & Arnott) Wawra: Itin. Princ. S. Coburgi 1: 17. (1883)
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