Horkelia fusca var. pseudocapitata |
Horkelia fusca var. fusca |
|
---|---|---|
intermountain horkelia, tawny horkelia |
horkelia, pinewoods horkelia, tawny horkelia |
|
Stems | 3–6 dm. |
3–4(–5) dm. |
Basal leaves | green, 10–35(–40) cm; leaflets 4–6(or 7) per side, narrowly to broadly obovate, 10–30(–35) × (5–)10–20(–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate. |
green, (4–)8–20 cm; leaflets 6–10 per side, cuneate-obovate, 5–18(–20) × 4–15 mm, 1/2 to nearly as wide as long, divided ± 1/2 to midrib into 5–8 teeth or lobes, surfaces not obscured, ± sparsely short-villous. |
Cauline leaves | 1–4(or 5); leaflets of proximalmost 2 or 3(–5) per side. |
(2 or)3–5; leaflets of proximalmost 3–6 per side. |
Inflorescences | green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular hairs not or obscurely red-septate; bracts acute-lobed, not obscuring pedicels and flowers at maturity. |
reddish purple, congested to open, comprising less than 1/4 of stem, usually composed of 10–20-flowered glomerules, rarely flowers arranged individually, glandular hairs often ± red-septate; bracts acute- to acuminate-lobed, not obscuring pedicels and flowers at maturity. |
Flowers | epicalyx bractlets 2–3.5(–4) mm; hypanthium 2 × 2.5–4 mm; petals (3–)4–6 mm; filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm; styles 1–1.5 mm. |
epicalyx bractlets 1.5–3 mm; hypanthium 1.5–3 × 2.5–4 mm; petals 3–5 mm; filaments 0.5–1.3 mm, longer than wide, anthers 0.5–0.6 mm; styles 1–1.5 mm. |
Achenes | 1.6–1.8 mm. |
1.6–1.8 mm. |
2n | = 28. |
|
Horkelia fusca var. pseudocapitata |
Horkelia fusca var. fusca |
|
Phenology | Flowering summer. | Flowering summer. |
Habitat | Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil | Moist meadows, in conifer woodlands, mainly on volcanic soil |
Elevation | 900–2300 m (3000–7500 ft) | 100–1500 m (300–4900 ft) |
Distribution |
CA; ID; NV; OR |
OR; WA |
Discussion | The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007). As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variety fusca occurs in the Cascade Ranges on both sides of the Columbia River Gorge, primarily around Mount Hood, Oregon, and Mount Adams, Washington. Pending further analysis, all collections of the species from the Cascade Range in Washington are treated here as var. fusca, including some with leaflets and/or petals in the size range of var. parviflora. Plants from lower elevations tend to have greener, more openly branched inflorescences and narrower, more finely divided leaflets; this expression provides the type of Horkelia tenuisecta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 262. | FNA vol. 9, p. 260. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | H. pseudocapitata, H. fusca subsp. pseudocapitata, Potentilla douglasii var. pseudocapitata | H. tenuisecta, Potentilla douglasii var. tenuisecta |
Name authority | (Rydberg ex Howell) M. Peck: Man. Pl. Oregon, 398. (1941) | unknown |
Web links |