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Photo is of parent taxon

intermountain horkelia, tawny horkelia

Photo is of parent taxon

Diamond Lake horkelia, pinewoods horkelia

Stems

3–6 dm.

(1–)1.5–3.5(–4) dm.

Basal leaves

green, 10–35(–40) cm;

leaflets 4–6(or 7) per side, narrowly to broadly obovate, 10–30(–35) × (5–)10–20(–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate.

green, 4–9(–12) cm;

leaflets 4–8 per side, cuneate-obovate, 5–12 × 2–8 mm, ± 1/2 as wide as long, divided ± 1/2+ to midrib into 3–9 teeth or lobes, surfaces not obscured, sparsely short-villous to glabrate.

Cauline leaves

1–4(or 5);

leaflets of proximalmost 2 or 3(–5) per side.

2–4(–6);

leaflets of proximalmost 2–5 per side.

Inflorescences

green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular hairs not or obscurely red-septate;

bracts acute-lobed, not obscuring pedicels and flowers at maturity.

green, open, comprising 1/3–1/2 of stem, flowers usually arranged individually, glandular hairs sometimes red-septate;

bracts acuminate-lobed, not obscuring pedicels and flowers at maturity.

Flowers

epicalyx bractlets 2–3.5(–4) mm;

hypanthium 2 × 2.5–4 mm;

petals (3–)4–6 mm;

filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm;

styles 1–1.5 mm.

epicalyx bractlets 1–2 mm;

hypanthium 1–2 × 2.5–3.5 mm;

petals 2–4(–5) mm;

filaments 0.5–1 mm, ± as wide as long, anthers 0.5 mm;

styles 1 mm.

Achenes

1.6–1.8 mm.

1.2–1.5 mm.

2n

= 28.

Horkelia fusca var. pseudocapitata

Horkelia fusca var. filicoides

Phenology Flowering summer. Flowering summer.
Habitat Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil Open conifer woodlands, mainly on volcanic soil
Elevation 900–2300 m (3000–7500 ft) 800–1600 m (2600–5200 ft)
Distribution
from FNA
CA; ID; NV; OR
from FNA
OR
[BONAP county map]
Discussion

The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007).

As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variety filicoides is known from the southern Cascade Range of southwestern Oregon, primarily in open lodgepole pine forests north and west of Crater Lake. This is the only variety in which the flowers are usually arranged individually rather than clustered into glomerules, at least in fully expanded inflorescences; some populations out of the core range have more congested inflorescences and are transitional to var. parviflora in this regard.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 262. FNA vol. 9, p. 260.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca
Sibling taxa
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. filicoides, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. tenella
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. pseudocapitata, H. fusca var. tenella
Synonyms H. pseudocapitata, H. fusca subsp. pseudocapitata, Potentilla douglasii var. pseudocapitata Potentilla douglasii var. filicoides, H. fusca subsp. filicoides
Name authority (Rydberg ex Howell) M. Peck: Man. Pl. Oregon, 398. (1941) (Crum) M. Peck: Man. Pl. Oregon, 399. (1941)
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