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Diamond Lake horkelia, pinewoods horkelia

Habit Herbs, shrubs, or subshrubs.
Stems

(1–)1.5–3.5(–4) dm.

Leaves

alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;

stipules present, rarely absent.

Basal leaves

green, 4–9(–12) cm;

leaflets 4–8 per side, cuneate-obovate, 5–12 × 2–8 mm, ± 1/2 as wide as long, divided ± 1/2+ to midrib into 3–9 teeth or lobes, surfaces not obscured, sparsely short-villous to glabrate.

Cauline leaves

2–4(–6);

leaflets of proximalmost 2–5 per side.

Inflorescences

green, open, comprising 1/3–1/2 of stem, flowers usually arranged individually, glandular hairs sometimes red-septate;

bracts acuminate-lobed, not obscuring pedicels and flowers at maturity.

Flowers

epicalyx bractlets 1–2 mm;

hypanthium 1–2 × 2.5–3.5 mm;

petals 2–4(–5) mm;

filaments 0.5–1 mm, ± as wide as long, anthers 0.5 mm;

styles 1 mm.

torus usually enlarged, sometimes small or absent;

carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);

ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).

Fruits

achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;

styles persistent or deciduous, not elongate (elongate but not plumose in Geum).

Achenes

1.2–1.5 mm.

x

= 7(8).

Horkelia fusca var. filicoides

Rosaceae subfam. rosoideae

Phenology Flowering summer.
Habitat Open conifer woodlands, mainly on volcanic soil
Elevation 800–1600 m (2600–5200 ft)
Distribution
from FNA
OR
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
Discussion

Variety filicoides is known from the southern Cascade Range of southwestern Oregon, primarily in open lodgepole pine forests north and west of Crater Lake. This is the only variety in which the flowers are usually arranged individually rather than clustered into glomerules, at least in fully expanded inflorescences; some populations out of the core range have more congested inflorescences and are transitional to var. parviflora in this regard.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.

Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 260. FNA vol. 9, p. 23. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca Rosaceae
Sibling taxa
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. pseudocapitata, H. fusca var. tenella
Subordinate taxa
Synonyms Potentilla douglasii var. filicoides, H. fusca subsp. filicoides
Name authority (Crum) M. Peck: Man. Pl. Oregon, 399. (1941) Arnott: Botany, 107. (1832)
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