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Diamond Lake horkelia, pinewoods horkelia

Habit Plants forming tufts or open mats, green or reddish, rarely grayish, conspicuously glandular, resinously aromatic.
Stems

(1–)1.5–3.5(–4) dm.

ascending to erect, (0.6–)1–6(–9) dm.

Basal leaves

green, 4–9(–12) cm;

leaflets 4–8 per side, cuneate-obovate, 5–12 × 2–8 mm, ± 1/2 as wide as long, divided ± 1/2+ to midrib into 3–9 teeth or lobes, surfaces not obscured, sparsely short-villous to glabrate.

planar;

stipules entire;

leaflets 3–8(–15) per side, separate, sometimes ± overlapping, divided 1/5–3/4+ to midrib into (3–)5–15 teeth or lobes not restricted to apex.

Cauline leaves

2–4(–6);

leaflets of proximalmost 2–5 per side.

Inflorescences

green, open, comprising 1/3–1/2 of stem, flowers usually arranged individually, glandular hairs sometimes red-septate;

bracts acuminate-lobed, not obscuring pedicels and flowers at maturity.

open to ± congested, flowers usually arranged in ± capitate glomerules, arranged individually in H. fusca var. filicoides.

Pedicels

remaining ± straight, outermost sometimes ± reflexed in congested inflorescences, 1–3(–10) mm.

Flowers

epicalyx bractlets 1–2 mm;

hypanthium 1–2 × 2.5–3.5 mm;

petals 2–4(–5) mm;

filaments 0.5–1 mm, ± as wide as long, anthers 0.5 mm;

styles 1 mm.

epicalyx bractlets linear, 0.2–0.3(–0.5) mm wide, entire;

hypanthium interior glabrous;

sepals acute to acuminate;

petals white to pale pink, often veined with pink to rose, ± oblanceolate to cuneate, apex emarginate to truncate or rounded;

filaments white, glabrous, anthers longer than wide;

carpels 10–25.

Achenes

1.2–1.5 mm.

1–1.8 mm, smooth.

Horkelia fusca var. filicoides

Horkelia sect. Capitatae

Phenology Flowering summer.
Habitat Open conifer woodlands, mainly on volcanic soil
Elevation 800–1600 m (2600–5200 ft)
Distribution
from FNA
OR
[BONAP county map]
w United States
Discussion

Variety filicoides is known from the southern Cascade Range of southwestern Oregon, primarily in open lodgepole pine forests north and west of Crater Lake. This is the only variety in which the flowers are usually arranged individually rather than clustered into glomerules, at least in fully expanded inflorescences; some populations out of the core range have more congested inflorescences and are transitional to var. parviflora in this regard.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 1.

Rydberg recognized seven species in his group Capitatae, treated here as intergrading variation within a single species. Plants of the section share the characteristic Horkelia odor, glandularity, and planar leaves of sect. Horkelia, but differ in the combination of relatively small, short-pedicelled flowers that (except for var. filicoides) are most commonly aggregated into one or more capitate, purple-suffused glomerules, with linear epicalyx bractlets, oblanceolate-cuneate petals that are often pink-tinged, and relatively short, broad filaments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 260. FNA vol. 9, p. 259.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Capitatae > Horkelia fusca Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia
Sibling taxa
H. fusca var. brownii, H. fusca var. capitata, H. fusca var. fusca, H. fusca var. parviflora, H. fusca var. pseudocapitata, H. fusca var. tenella
Subordinate taxa
Synonyms Potentilla douglasii var. filicoides, H. fusca subsp. filicoides H. unranked Capitatae, Potentilla section Capitatae
Name authority (Crum) M. Peck: Man. Pl. Oregon, 399. (1941) (Rydberg) O. Stevens: in N. L. Britton et al., N. Amer. Fl. 22(7): 7. (1959)
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