Horkelia fusca var. filicoides
Horkelia fusca var. brownii
Diamond Lake horkelia, pinewoods horkelia
Shasta horkelia, tawny horkelia
(1–)1.5–3.5(–4) dm.
1–4.5 dm.
green, 4–9(–12) cm;
leaflets 4–8 per side, cuneate-obovate, 5–12 × 2–8 mm, ± 1/2 as wide as long, divided ± 1/2+ to midrib into 3–9 teeth or lobes, surfaces not obscured, sparsely short-villous to glabrate.
grayish, 4–12(–18) cm;
leaflets 3–7 per side, narrowly to broadly cuneate, 5–15(–17) × 2–8(–12) mm, usually 1/3–1/2 as wide as long, divided 1/4–1/2 to midrib into 4–6 teeth, surfaces often obscured, ± densely short-villous.
2–4(–6);
leaflets of proximalmost 2–5 per side.
2–4;
leaflets of proximalmost 2–4 per side.
green, open, comprising 1/3–1/2 of stem, flowers usually arranged individually, glandular hairs sometimes red-septate;
bracts acuminate-lobed, not obscuring pedicels and flowers at maturity.
grayish or purplish, usually ± congested, sometimes more open, usually comprising less than 1/6 of stem, composed of 5–20(–40)-flowered glomerules, glandular hairs usually not red-septate;
bracts acuminate-lobed, often ± obscuring pedicels and flowers even at maturity.
epicalyx bractlets 1–2 mm;
hypanthium 1–2 × 2.5–3.5 mm;
petals 2–4(–5) mm;
filaments 0.5–1 mm, ± as wide as long, anthers 0.5 mm;
styles 1 mm.
epicalyx bractlets 1.5–2.5 mm;
hypanthium 1.5–2.5 × 2.5–4 mm;
petals 2–3(–4.5) mm;
filaments (0.2–)0.6–1 mm, usually longer than wide, anthers 0.3–0.5 mm;
styles 1 mm.
1.2–1.5 mm.
1–1.2 mm.
= 28.
Horkelia fusca var. filicoides
Horkelia fusca var. brownii
Variety filicoides is known from the southern Cascade Range of southwestern Oregon, primarily in open lodgepole pine forests north and west of Crater Lake. This is the only variety in which the flowers are usually arranged individually rather than clustered into glomerules, at least in fully expanded inflorescences; some populations out of the core range have more congested inflorescences and are transitional to var. parviflora in this regard.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Variety brownii here takes the place of var./subsp. pseudocapitata as used in D. D. Keck (1938) and most floras; this variant was included within var. parviflora by B. Ertter (1993d). The diagnostic feature is the ash gray leaflets, which also tend to be more narrowly cuneate than in other varieties. The variety occurs from Mount Shasta in Siskiyou County east to southwestern Modoc County and southeast to Sierra Valley in Plumas and Sierra counties, California, skirting the range of var. tenella. Morphologically comparable plants occur as far south as the California-Nevada border on the south side of Lake Tahoe, and at least one collection from Nevada (Reno, Hillman s.n., POM) is closest to this variety. Populations from the southeastern end of the range tend to be larger and less ash gray than those in the vicinity of Mount Shasta. Most references to var. brownii (as var./subsp. pseudocapitata misapplied) from Oregon are probably best attributed to the polymorphic var. parviflora, but at least one collection from Klamath Lake (Newberry s.n., GH) fits well in var. brownii.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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