Flora of North America species comparison

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Horkelia daucifolia var. indicta

Horkelia sect. Tridentatae
Photo is of parent taxon

Jepson's horkelia
Habit Plants rosette-forming to tufted. Plants forming rosettes or tufts, sometimes loose mats, green to grayish or silvery, inconspicuously to moderately glandular, not resinously aromatic.
Stems

decumbent to erect, 0.5–5 dm.
Basal leaves

green, 7–11 cm;

leaflets 5–10(–15) mm, lobes (2–)5–15, linear, 0.4–1 mm wide.

planar to ± cylindric;

stipules entire, forked, or pinnately divided into linear to filiform lobes;

leaflets 2–20 per side, separate to ± overlapping, divided either 1/2–3/4+ to midrib into (0–)2–15 ultimate lobes or teeth not restricted to apex, or 1/10–1/4 or less to midrib into (0–)3(–5) teeth restricted to apex.
Inflorescences

open to ± congested, flowers arranged individually, in corymbiform clusters, and/or in capitate or non-capitate glomerules.
Pedicels

5–10(–20) mm, not puberulent, minutely glandular.

remaining straight, outermost sometimes ± reflexed in congested inflorescences, 1–10(–20) mm.
Flowers

petals broadly obovate to obcordate, 5–7.5 × 4–6 mm;

filaments 2–3 mm.

epicalyx bractlets linear to lanceolate or narrowly elliptic, 0.2–0.5(–0.8) mm wide, entire;

hypanthium interior usually glabrous, sometimes pilose;

sepals obtuse to acute to slightly acuminate;

petals white to pink or cream, sometimes drying yellowish, linear or cuneate to obovate or obcordate, apex usually acute to rounded to emarginate, sometimes truncate, rarely mucronate;

filaments white, glabrous, anthers longer or shorter than wide;

carpels 2–15(–20).
Achenes

1.5–3 mm, smooth or rugose.

Horkelia daucifolia var. indicta

Horkelia sect. Tridentatae
Phenology Flowering summer.
Habitat Vernally saturated openings in oak-conifer woodlands, mainly on clay soil
Elevation 200–700 m (700–2300 ft)
Distribution
from FNA
CA
[BONAP county map]
w United States
Discussion

Of conservation concern.

Variety indicta was known only from the type in Tehama County until rediscovered in 1990 in western Shasta County, confirming that this is a distinct variety in the forested hills rimming the northern Sacramento Valley (B. Ertter and J. L. Reveal 2007). One other historic collection of Horkelia daucifolia from Shasta County (Montgomery Creek, Smith s.n., CAS) is more similar to var. daucifolia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 5 (5 in the flora).

P. A. Rydberg (1908c) assigned the species here composing this section to three of his unranked groups: Hirsutae, Sericatae (also including Horkelia hispidula), and Tridentatae, validated inadvertently as sections by O. A. Stevens in the index to volume 22 of North American Flora. In addition to lacking the characteristic Horkelia odor, plants are often rosette-forming from simple caudices; stipules are sometimes highly divided; and leaflets are either few-toothed or deeply lobed. The section is centered in the Siskiyou-Klamath region of northwestern California and southwestern Oregon, with some affinity for ultramafic substrates. Horkelia tridentata extends south in the Sierra Nevada; H. congesta is a component of the Willamette Valley flora. Much of the variation in the section is highly localized, with a full series of intergrading morphologies, indicative of a relatively recent and active radiation. More segregates are recognized here than in some recent floras (M. E. Peck 1941; P. A. Munz 1959; B. Ertter 1993d), and additional variants are likely to warrant taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Leaflets 2–6 per side, separate to slightly overlapping, divided into (0–)3(–5) teeth restricted to apex; inflorescences: flowers usually arranged in ± capitate glomerules, sometimes in corymbiform clusters
→ 2
1. Leaflets 5–20 per side, ± overlapping, divided into (0–)2–15 linear to elliptic or oblanceolate lobes or teeth not restricted to apex; inflorescences: flowers arranged individually, in corymbiform clusters, and/or in usually non-capitate glomerules
→ 3
2. Petals ± obovate, (1.5–)2–5 mm wide; stems: glands dense distally; Oregon, barely in nw California.
H. congesta
2. Petals linear to broadly oblanceolate, 0.3–1.5 mm wide; stems: glands absent or sparse distally; California, barely in sw Oregon.
H. tridentata
3. Leaflets 5–10 per side, 5–15(–25) mm, divided 3/4+ to midrib into (0–)2–15 lobes; stems: hairs 2–3 mm proximally; inflorescences open to ± congested, flowers arranged individually or in ± corymbiform clusters; petals white to cream, often drying yellowish.
H. daucifolia
3. Leaflets (8–)10–20 per side, 2–10(–15) mm, divided ± 1/2–3/4 to midrib into (0–)2–4 lobes; stems: hairs 1 mm proximally; inflorescences open, flowers arranged individually and/or in non-capitate glomerules; petals white to pink or red-veined
→ 4
4. Plants silvery; basal leaves densely sericeous, 3–10 cm, stipules usually entire or forked, rarely pinnately divided.
H. sericata
4. Plants ± green; basal leaves usually villous to pilose, often glabrate, 5–15 cm, stipules deeply 2-lobed or pinnately divided into 3–5 lobes.
H. howellii
Source FNA vol. 9, p. 267. FNA vol. 9, p. 264.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Tridentatae > Horkelia daucifolia Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia
Sibling taxa
H. daucifolia var. caruifolia, H. daucifolia var. daucifolia
Subordinate taxa
H. congesta, H. daucifolia, H. howellii, H. sericata, H. tridentata
Synonyms Potentilla daucifolia var. indicta H. unranked Tridentatae, H. unranked Ambiguae, H. unranked Hirsutae, H. section Hirsutae, H. unranked Sericatae, H. section Sericatae
Name authority (Jepson) Ertter & Reveal: Novon 17: 319. (2007) (Rydberg) O. Stevens: in N. L. Britton et al., N. Amer. Fl. 22(7): 7. (1959)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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