Horkelia cuneata var. sericea
Horkelia cuneata
coast horkelia, Kellogg's horkelia, wedgeleaf horkelia
wedge leaf horkelia
decumbent to ascending, 2–5(–7.5) dm.
decumbent to erect, (1–)2–6(–7.5) dm, hairs ascending to appressed, sometimes spreading.
planar, (5–)10–30 × 1.5–4(–5) cm;
stipules entire or basally lobed;
leaflets 5–12 per side, separate to slightly overlapping, ± elliptic to obovate or oblanceolate, 5–25(–30) × 5–15(–25) mm, ± 1/2 to nearly as wide as long, divided less than 1/3 to midrib into (5–)10–15 oblanceolate to obovate teeth, sparsely pilose to glabrate or sericeous.
1–3(–5).
5–10 per side, obovate, (5–)10–25(–30) mm, not distinctly pinnately veined.
usually ± congested;
most flowers arranged in glomerules.
± open to congested, flowers arranged individually and/or in glomerules.
1–3 mm, proximalmost to 12 mm.
1–30(–40) mm.
hypanthium interior rim densely pilose;
petals oblanceolate, 1.5–3 mm wide;
filaments 1–3 × 0.5–1 mm.
12–17 mm diam.;
epicalyx bractlets lanceolate to ovate, 3–5 × 1.5–3 mm, 2/3–3/4 length of sepals, entire;
hypanthium 1.5–2 × 4–7 mm, less than 1/2 as deep as wide, interior glabrous or pilose in a narrow ring;
sepals ± reflexed to spreading, lanceolate, 4–6.5 mm;
petals oblanceolate to narrowly obovate, 4–8 × 1.5–4 mm, apex obtuse to rounded;
filaments 1–3 × 0.5–2 mm, anthers 0.7–1.1 mm;
carpels (30–)40–60(–80);
styles 2–3 mm.
brown, 1.5–1.8 mm, smooth or minutely rugose.
= 28.
Horkelia cuneata var. sericea
Horkelia cuneata
Of conservation concern.
Historic populations of var. sericea occurred along the immediate coast from Alameda and Marin counties south at least to Santa Barbara County, with some littoral collections of Horkelia cuneata from Los Angeles and San Diego counties having some sericea characteristics. Reports from farther north are all based on misidentifications, for example, of H. californica in Sonoma County. The most distinctive specimens are from the northern populations in Alameda, Marin, San Francisco, and San Mateo counties, all of which apparently no longer exist. Of the recently confirmed extant populations, those that come closest to justifying continued recognition of var. sericea are in Monterey, San Luis Obispo, and Santa Cruz counties.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 3 (3 in the flora).
As noted by D. D. Keck (1938) and B. Ertter (1997b), recognition of subunits within Horkelia cuneata is complicated by the abundance of intermediates, to the extent that a complete transition exists between the decumbent, densely hairy coastal extreme with a congested inflorescence, represented by var. sericea, and the erect, green-glandular interior extreme with an open inflorescence, represented by var. puberula. The traditional division into three infraspecific units is maintained here to emphasize the distinctiveness of the extremes, which would qualify as specifically distinct were they not ends of a continuum. The extremes are furthermore threatened by ongoing decimation of suitable habitats at the northern (San Francisco Bay Area) and southern (greater Los Angeles) ends of the species range.
Determining the correct name of Horkelia cuneata if treated as a species of Potentilla is complicated. Potentilla cuneata (Lindley) Baillon ex Munz & I. M. Johnston (1925) and P. puberula Greene are both later homonyms, while P. multijuga Lehmann has been conserved with a conserved type to maintain established usage for an unrelated species of Potentilla (B. Ertter and J. L. Reveal 2008). J. T. Howell (1945) adopted P. lindleyi; however, as noted by R. F. Hoover (1966), P. kelloggii has priority at species rank by a matter of months. The latter is accordingly the correct name in Potentilla for H. cuneata.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Plants: eglandular hairs sparse or absent, glandular hairs evident; hypanthium interior rim usually glabrous, sometimes sparsely pilose; pedicels (2–)5–10 mm, proximalmost to 40 mm; stems erect. | var. puberula |
1. Plants: eglandular hairs moderately abundant to dense, glandular hairs evident or obscured; hypanthium interior rim ± or densely pilose; pedicels 1–5 mm, proximalmost to 15 mm; stems decumbent to erect | → 2 |
2. Plants ± green, eglandular hairs moderately abundant, spreading to ascending, glandular hairs usually evident; stems ascending to erect. | var. cuneata |
2. Plants grayish-sericeous, eglandular hairs dense, ascending to appressed, glandular hairs obscured; stems decumbent to ascending. | var. sericea |
CA
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