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mesa horkelia, wedgeleaf horkelia

Habit Plants green; eglandular hairs sparse or absent, spreading, glandular hairs not obscured. Herbs, shrubs, or subshrubs.
Stems

erect, 3–6 dm.

Leaves

alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;

stipules present, rarely absent.

Leaflets

5–12 per side, elliptic to obovate, 5–25 mm, distinctly pinnately veined.

Inflorescences

open, most flowers arranged individually.

Pedicels

(2–)5–10 mm, proximalmost to 40 mm.

Flowers

hypanthium interior rim usually glabrous, sometimes sparsely pilose;

petals oblanceolate to narrowly obovate, 2–4 mm wide;

filaments 1–2.5 × 0.5–2 mm.

torus usually enlarged, sometimes small or absent;

carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);

ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).

Fruits

achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;

styles persistent or deciduous, not elongate (elongate but not plumose in Geum).

x

= 7(8).

Horkelia cuneata var. puberula

Rosaceae subfam. rosoideae

Phenology Flowering spring–summer.
Habitat Coastal ranges away from the immediate coast, sandy to gravelly soil, coastal sage and chaparral communities
Elevation 70–900 m (200–3000 ft)
Distribution
from FNA
CA
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
Discussion

Of conservation concern.

At its most distinctive, var. puberula is represented by historic collections from the hills and plains of Los Angeles, western Riverside, extreme southwestern San Bernardino, and northwestern San Diego counties; these populations have now largely been destroyed by urbanization. Scattered populations still exist within this core area, to what extent and with what level of ensured continuity remains uncertain. In addition, multiple populations of Horkelia cuneata from as far north and west as San Luis Obispo County have characteristics of var. puberula; these tend to intergrade with var. cuneata, as do some historic collections from the core area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.

Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 255. FNA vol. 9, p. 23. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Horkelia > sect. Horkelia > Horkelia cuneata Rosaceae
Sibling taxa
H. cuneata var. cuneata, H. cuneata var. sericea
Subordinate taxa
Synonyms H. puberula, H. cuneata subsp. puberula, P. kelloggii var. puberula, P. lindleyi var. puberula
Name authority (Rydberg) Ertter & Reveal: Novon 17: 318. (2007) Arnott: Botany, 107. (1832)
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