Horkelia cuneata var. puberula
Horkelia sect. Horkelia
mesa horkelia, wedgeleaf horkelia
erect, 3–6 dm.
decumbent or ascending to erect, (0.5–)1–10(–12) dm.
usually planar, sometimes ± cylindric;
stipules usually entire, sometimes basally lobed;
leaflets (1–)3–16(–20) per side, separate to overlapping, divided ± 1/6–3/4+ to midrib into 3–30(–60) teeth or lobes not restricted to apex.
5–12 per side, elliptic to obovate, 5–25 mm, distinctly pinnately veined.
open, most flowers arranged individually.
open to congested, flowers arranged individually, in usually non-capitate glomerules, and/or in corymbiform clusters.
(2–)5–10 mm, proximalmost to 40 mm.
remaining straight, 1–30(–40) mm.
hypanthium interior rim usually glabrous, sometimes sparsely pilose;
petals oblanceolate to narrowly obovate, 2–4 mm wide;
filaments 1–2.5 × 0.5–2 mm.
epicalyx bractlets narrowly elliptic-lanceolate to broadly ovate, 0.5–3 mm wide, usually entire, sometimes toothed;
hypanthium interior pilose or glabrous;
sepals acute;
petals white, oblong-oblanceolate to round, apex obtuse to truncate to emarginate;
filaments white, glabrous, anthers longer than wide;
carpels 10–200(–220).
0.8–2 mm, usually smooth or slightly rugose, sometimes merely roughened.
Horkelia cuneata var. puberula
Horkelia sect. Horkelia
Of conservation concern.
At its most distinctive, var. puberula is represented by historic collections from the hills and plains of Los Angeles, western Riverside, extreme southwestern San Bernardino, and northwestern San Diego counties; these populations have now largely been destroyed by urbanization. Scattered populations still exist within this core area, to what extent and with what level of ensured continuity remains uncertain. In addition, multiple populations of Horkelia cuneata from as far north and west as San Luis Obispo County have characteristics of var. puberula; these tend to intergrade with var. cuneata, as do some historic collections from the core area.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species 9 (9 in the flora).
Section Horkelia encompasses the species that are most commonly encountered in heavily populated areas of California. Plants are notably glandular-viscid (unless obscured by dense vestiture) and have a distinctive resinous odor.
Previous revisions (for example, P. A. Rydberg 1908c; D. D. Keck 1938) have placed Horkelia frondosa (here treated as H. californica var. frondosa) at the beginning, implying that this is the least derived expression within the genus. Such an assumption is based on its gross resemblance to sympatric members of Drymocallis; molecular evidence (T. Eriksson et al. 1998; M. Lundberg et al. 2009; C. Dobeš and J. Paule 2010) confirms that this similarity is superficial. If, as speculated above, species composing sect. Hispidulae are relicts of the original radiation, then H. californica var. frondosa is actually one of the more derived members of the genus. Species within sect. Horkelia have been ordered here according to that interpretation.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaflets 1–3 per side, sparsely short-pilose to glabrate, (15–)20–30-toothed 1/6–1/4 to midrib; petals obovate to round, (3–)4–7 mm wide. | H. truncata |
1. Leaflets (3–)5–16(–20) per side, hairy, 3–15(–30)-toothed or -lobed (collectively 10–60-toothed in H. californica) 1/6–3/4+ to midrib; petals narrowly obovate or oblanceolate to elliptic or oblong, 1–3(–4) mm wide | → 2 |
2. Hypanthia 3–5.5 mm, 1/2 to nearly as deep as wide; epicalyx bractlets 4–6(–8) mm, ± equal to sepals, entire or toothed. | H. californica |
2. Hypanthia 1–2(–3) mm, less than 1/2 as deep as wide (except H. clevelandii); epicalyx bractlets 1–4(–5) mm, shorter than sepals, entire | → 3 |
3. Leaflets ± elliptic to obovate, divided less than 1/3 to midrib into (5–)10–15 teeth; epicalyx bractlets 1.5–3 mm wide. | H. cuneata |
3. Leaflets cuneate or flabellate to ovate or nearly round, divided (1/5–)1/3–3/4+ to midrib into 3–10(–30 in H. yadonii) teeth or lobes; epicalyx bractlets 0.5–2(–2.5) mm wide | → 4 |
4. Stems ± decumbent to ascending, 1–3.5(–4.5) dm; leaflets 5–10(–12) per side, cuneate, ± overlapping; inflorescences usually congested; plants matted; coastal. | H. marinensis |
4. Stems ascending to erect, (0.5–)1–7 dm; leaflets 6–16(–20) per side, cuneate to flabellate or nearly round, separate to ± overlapping; inflorescences open to ± congested; plants tufted to matted; coastal or interior | → 5 |
5. Epicalyx bractlets 1–2(–2.5) mm wide, lanceolate to ovate; anthers 0.8–1.2 mm; hypanthia 3–6.5 mm diam. | H. yadonii |
5. Epicalyx bractlets 0.5–1(–1.5) mm wide, narrowly elliptic to broadly lanceolate; anthers 0.4–1 mm; hypanthia 2–4.5(–6) mm diam | → 6 |
6. Stems: hairs ascending to appressed; plants usually grayish to grayish green; hypanthium interior pilose | → 7 |
6. Stems: hairs ± spreading; plants green to grayish; hypanthium interior glabrous or pilose | → 8 |
7. Carpels 20–50(–120); styles 2–4 mm; basal leaves (4–)8–20(–30) cm; Transverse Ranges, s California. | H. rydbergii |
7. Carpels 10–20(–27); styles (1–)1.5–2 mm; basal leaves 3–8(–9) cm; Coast Ranges, n California. | H. bolanderi |
8. Leaflets divided 1/2–3/4+ to midrib into linear to narrowly oblanceolate or narrowly elliptic lobes; hypanthium interior pilose; n Coast Ranges, California. | H. tenuiloba |
8. Leaflets divided 1/3 to midrib into acute to obtuse teeth; hypanthium interior glabrous; Peninsular Ranges, s California. | H. clevelandii |
CA
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