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bulbous barley, charming barley (ssp. leporinum), foxtail barley, hare barley, mouse barley, mouse barley (ssp. murinum), seagreen barley (ssp. glaucum), smooth barley, wall barley

barley

Habit Plants annual; loosely tufted. Plants summer or winter annuals or perennials; cespitose, sometimes shortly rhizomatous.
Culms

to 110 cm, usually erect, sometimes almost prostrate;

nodes glabrous.

to 135(150) cm, erect, geniculate, or decumbent;

nodes glabrous or pubescent.

Sheaths

open, pubescent or glabrous;

auricles present or absent;

ligules hyaline, truncate, erose;

blades flat to more or less involute, more or less pubescent on both sides.

Inflorescences

usually spikelike racemes, sometimes spikes, all customarily called spikes, with 3 spikelets at each node, central spikelets usually sessile, sometimes pedicellate, pedicels to 2 mm, lateral spikelets usually pedicellate, pedicels curved or straight, sometimes all 3 spikelets sessile in cultivated plants;

disarticulation usually in the rachises, the spikelets falling in triplets, cultivated forms generally not disarticulating.

Spikes

3-8 cm long, 7-16 mm wide, pale green to distinctly reddish, especially the awns;

rachises disarticulating at maturity.

Spikelets

with 1 floret;

glumes awnlike, usually exceeding the floret.

Caryopses

usually tightly enclosed in the lemma and palea at maturity.

Lower

sheaths often completely surrounding the culms, glabrous or somewhat pilose;

ligules 1-4 mm;

auricles to 8 mm, well developed even on the upper leaves;

blades to 28 cm, usually flat, occasionally with involute margins, glabrous or sparsely pilose, sometimes scabrous.

Central

spikelets sessile, florets sessile or pedicellate, pedicels to 2 mm;

glumes 11-25 mm long, 0.8-1.8 mm wide, flattened, margins usually distinctly ciliate;

lemmas 8-14 mm long, to 2 mm wide, more or less smooth, awned, awns 20-40 mm;

lodicules glabrous or with 1+ cilia;

anthers 0.2-3.2 mm, gray to yellow, sometimes with purple spots.

spikelets bisexual;

florets sessile;

rachillas prolonged beyond the floret;

lemmas ovate, glabrous to pubescent, 5-veined, usually awned, rarely unawned;

paleas almost equal to the lemmas, narrowly ovate, keeled;

lodicules 2, broadly lanceolate, margins ciliate;

anthers 3, usually yellowish.

Lateral

spikelets staminate, floret sessile;

glumes flattened, margins ciliate;

lemmas 8-15 mm, awned, awns 20-50 mm;

paleas 8-15 mm;

rachillas 2.5-6.5 mm, slender or gibbous, yellow.

spikelets usually sterile or staminate, often bisexual in cultivated forms;

florets pedicellate, usually reduced;

lemmas awned or unawned.

2n

= 14, 28, 42.

= 14, 28, 42.

Hordeum murinum

Hordeum

Distribution
from FNA
AL; AZ; CA; CO; CT; DC; DE; GA; ID; MA; MD; ME; MT; NC; NJ; NM; NV; NY; OK; OR; PA; SC; TX; UT; VA; WA; WY; HI; AB; BC
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from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
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Discussion

Hordeum murinum is native to Eurasia, where it is a common weed in areas of human disturbance. It is thought to have originated around seasides, sandy riverbanks, and animal watering holes. It is now an established weed in the southwestern Flora region and other scattered locations. The records in Alaska are from the Anchorage area. Prostrate plants are associated with grazing. Three subspecies are recognized.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hordeum is a genus of 32 species that grow in temperate and adjacent subtropical areas, at elevations from 0-4500 m. The genus is native to Eurasia, the Americas, and Africa, and has been introduced to Australasia. The species are confined to rather moist habitats, even on saline soils. The annual species occupy seasonally moist habitats that cannot sustain a continuous grass cover.

Some species of Hordeum, such as H. marinum and H. murinum, are cosmopolitan weeds. Hordeum vulgare is widely cultivated for feed, malt, and flour. Archeological records suggest that Hordeum and Triticum were two of the earliest domesticated crops.

Eleven species of Hordeum grow in the Flora region: six are native, three are established weeds, and two are cultivated and occasionally persist as weeds. Hordeum secalinum has been reported from the Flora region, but the reports are based on misidentifications.

Four different haplomes are present in Hordeum. Hordeum vulgare and H. bulbosum have the I genome (often called the H genome by plant breeders), North American diploid species are based on the H genome, diploid H. marinum on the X genome, and diploids in the H. murinum group on the Y genome. Relationships among the polyploid taxa are complex (Jakob and Blattner 2006).

Spike measurements and lemma lengths, unless stated otherwise, do not include the awns.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spikelets sessile to subsessile; lemmas of the central florets subequal to those of the lateral florets, the awns longer than those of the lateral florets; paleas of the lateral florets almost glabrous
subsp. murinum
1. Central spikelets pedicellate; lemmas of the central florets from subequal to shorter than those of the lateral florets, the awns from shorter to longer than those of the lateral florets; paleas of the lateral florets scabrous to hairy.
→ 2
2. Lemmas of the central florets much shorter than those of the lateral florets; paleas of the lateral florets scabrous on the lower 1/2; anthers of the central and lateral florets similar in size
subsp. leporinum
2. Lemmas of the central florets about equal to those of the lateral florets; paleas of the lateral florets distinctly pilose on the lower 1/2; anthers of the central florets 0.2-0.6 mm long, those of the lateral florets 1.2-1.8 mm long
subsp. glaucum
1. Plants perennial.
→ 2
2. Culms usually with a bulbous swelling at the base; auricles to 5.5 mm long, well developed
H. bulbosum
2. Culms not bulbous-based; auricles absent or no more than 1 mm long.
→ 3
3. Glumes of the central spikelet flattened near the base
H. arizonicum
3. Glumes of the central spikelet usually setaceous throughout, rarely flattened near the base.
→ 4
4. Glumes 15-85 mm long, divergent to strongly divergent at maturity
H. jubatum
4. Glumes 7-19 mm long, divergent or not at maturity.
→ 5
5. Anthers of the central spikelet 0.8-4 mm long; auricles absent
H. brachyantherum
5. Anthers of the central spikelet 3.5-5 mm long; auricles present on the basal leaves
H. secalinum
1. Plants annual.
→ 6
6. Auricles to 8 mm long, well developed even on the upper leaves; lemmas of the lateral florets 6-15 mm long.
→ 7
7. Rachises disarticulating at maturity; glumes of the central spikelets ciliate; lemmas of the central florets to 2 mm wide, with awns 20-40 mm long; lateral spikelets staminate
H. murinum
7. Rachises usually not disarticulating at maturity; glumes of the central spikelets pubescent; lemmas of the central florets at least 3 mm wide, unawned or with awns 30-180 mm long; usually 1 or both lateral spikelets at a node seed-forming
H. vulgare
6. Auricles usually absent or to 0.3 mm long; lemmas of the lateral florets 1.7-8.5 mm long.
→ 8
8. Glumes bent, strongly divergent at maturity.
→ 9
9. Glumes of the central spikelets not flattened, (15)35-85 mm long
H. jubatum
9. Glumes of the central spikelets slightly flattened towards the base, 11-28 mm long
H. arizonicum
8. Glumes straight, ascending to slightly divergent at maturity.
→ 10
10. Lemmas of the lateral spikelets with awns 3-8 mm long
H. marinum
10. Lemmas of the lateral spikelets unawned or with awns no more than 3 mm long.
→ 11
11. Glumes of the central spikelets setaceous to slightly flattened near the base.
→ 12
12. Spikes 4-8 mm wide; lemmas of the central spikelets with awns 3-12 mm long; ligules 0.3-0.8 mm
H. depressum
12. Spikes 6-20 mm wide; lemmas of the central spikelets with awns 10-22 mm long; ligules 0.6-1.8 mm long
H. arizonicum
11. Glumes of the central spikelets distinctly flattened near the base.
→ 13
13. Lemmas of the lateral spikelets 1.7-4.4 mm long, usually unawned, rarely with awns to 1.2 mm long; sheaths with stripes of hairs
H. intercedens
13. Lemmas of the lateral spikelets 2.5-5.7 mm long, usually awned, with awns to 1.8 mm long; sheaths glabrous
H. pusillum
Source FNA vol. 24, p. 250. FNA vol. 24, p. 245. Author: Roland von Bothmer; Claus Badenf; Niels H. Jacobsen;.
Parent taxa Poaceae > subfam. Pooideae > tribe Triticeae > Hordeum Poaceae > subfam. Pooideae > tribe Triticeae
Sibling taxa
H. arizonicum, H. brachyantherum, H. bulbosum, H. depressum, H. intercedens, H. jubatum, H. marinum, H. pusillum, H. secalinum, H. vulgare
Subordinate taxa
H. murinum subsp. glaucum, H. murinum subsp. leporinum, H. murinum subsp. murinum
H. arizonicum, H. brachyantherum, H. bulbosum, H. depressum, H. intercedens, H. jubatum, H. marinum, H. murinum, H. pusillum, H. secalinum, H. vulgare
Name authority L. L.
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