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false spikeflower, hollisteria

false spikeflower, false spineflower

Habit Herbs, annual; taproots slender. Plants 0.3–0.8(–1) × (0.5–)0.8–3(–5) dm.
Stems

arising directly from the root, spreading to decumbent or prostrate, solid, not fistulose or disarticulating into ringlike segments, tomentose.

Leaves

deciduous (basal) or persistent (cauline), basal and cauline, alternate;

petiole indistinct at proximal node, absent at distal nodes;

blade oblanceolate (proximal) or elliptic to ovate (distal), margins entire, apex mucronate.

basal blades (1.5–)2–5(–6) × (0.2–)0.3–0.7(–0.9) cm;

cauline blades (0.5–)1–3.5 × (0.1–)0.3–0.8(–1) cm.

Inflorescences

terminal, cymose, distally uniparous due to suppression of secondaries, each node bearing a flower;

branches dichotomous, not brittle or disarticulating into segments, round, tomentose;

bracts 3, typically situated at base of a cauline leaf and hidden by it, connate basally, linear to linear-lanceolate, somewhat foliaceous or scalelike, awned, thinly pubescent to densely tomentose.

bracts dimorphic, lateral 2 obvious, 2–5 mm, thinly pubescent, central 1 obscured, 1–2(–3) mm, densely tomentose.

Peduncles

absent.

Flowers

1 per involucral cluster, sessile or pedicellate;

perianth yellowish, broadly campanulate when open, tubular when closed, densely tomentose abaxially;

tepals 6, connate proximally, monomorphic, mucronate apically;

stamens 6 or 9;

filaments basally adnate, glabrous;

anthers yellow, oval to oblong.

perianth 1.5–2 mm.

Achenes

included, brown to black, not winged, 3-gonous, glabrous.

1.7–2 mm.

Seeds

embryo curved.

Involucral

bracts obvious, in whorl of 3(–4), linear to lanceolate, awned.

bracts 2 mm, densely tomentose.

x

= 21.

2n

= 42.

Hollisteria

Hollisteria lanata

Phenology Flowering Mar–Jul.
Habitat Sandy to gravelly or clayey places, mixed grassland, desert scrub, chaparral communities, woodlands
Elevation 10-1000 m (0-3300 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 1.

Hollisteria is the only member of subtribe Hollisteriineae H. Gross. This view differs from that presented by J. L. Reveal and C. B. Hardham (1989), who included Chorizanthe and its relatives also in Hollisteriineae. In an unpublished thesis, Adrienne Russell (pers. comm.) has suggested that Hollisteria is more closely related to Eriogonum than to Chorizanthe. Reveal and Hardham noted that the transfer of Hollisteria from a place near Eriogonum to a position with Chorizanthe was done with “some trepidation.” There is no obvious place within Eriogonum subg. Ganysma to suggest as a point of origin for Hollisteria. Then, too, there is no place within Chorizanthe that one can comfortably suggest as such a point. By referring Hollisteria to its own subtribe, one acknowledges a relationship to Eriogonum, and recognizes that the single flower and base chromosome number of x = 21 are conditions unknown in the whole of Eriogonineae, although both features are common in Chorizanthineae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hollisteria lanata is encountered occasionally in the San Joaquin Valley and the adjacent coastal ranges and eastward in the Transverse Ranges to Kern and Tulare counties. Plants are only rarely locally common.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 444. Author: James L. Reveal. FNA vol. 5, p. 445.
Parent taxa Polygonaceae > subfam. Eriogonoideae Polygonaceae > subfam. Eriogonoideae > Hollisteria
Subordinate taxa
H. lanata
Synonyms Eriogonum section H. Eriogonum lanatum
Name authority S. Watson: Proc. Amer. Acad. Arts 14: 296. (1879) S. Watson: Proc. Amer. Acad. Arts 14: 296. (1879)
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