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leather-root

Loma Prieta, Loma Prieta hoita, Loma Prieta psoralea

Habit Herbs, perennial, unarmed; stoloniferous, with woody caudex or root.
Stems

erect or prostrate, often fistulose, usually pubescent and glandular-punctate;

cataphylls often present on proximalmost nodes.

erect, to 100 cm;

caudex woody;

proximalmost nodes without leaves or cataphylls.

Leaves

alternate, odd-pinnate;

stipules present;

petiolate;

leaflets 3, blade margins entire, surfaces pubescent, glandular.

stipules erect to reflexed, falcately lanceolate to caudate-elliptic, 7–16 mm, glandular, pubescent;

petiole 6–13.5 cm;

leaflet blades usually orbiculate or oblate, rarely ovate or lanceolate, 45–80 × 32–75 mm, base cordate or rounded, apex acute, surfaces pale and eglandular abaxially, dark brown to black-glandular and strigose adaxially.

Inflorescences

3–75+-flowered, axillary, pseudoracemes;

bracts soon or tardily deciduous.

long-ellipsoid, nodes 5–12, flowers 2 or 3 per node;

rachis 33–60 mm.

Peduncles

4.5–6.5 cm.

Flowers

papilionaceous;

calyx tubular-campanulate, lobes 5, not enlarging in fruit, becoming papery, lobes longer than tube;

corolla purple, or greenish tinged violet or purple, banner biauriculate, wings auriculate, keel always shorter than other petals, connate on apical margins, glabrous;

stamens 10, monadelphous early, vexillary stamen becoming distinct;

anthers in 2 series, proximal dorsifixed, distal basifixed;

ovule 1.

calyx 13–17 mm, dark brown- to black-glandular, hirsute to villose;

corolla purple, banner base greenish, 12–13 × 7.5–9 mm;

stamens 8.5–10.5 mm.

Fruits

legumes, included in calyx, longer than tube, stipitate, compressed, ovate to obovate indehiscent, eglandular, pubescent, with secondary, internal wall of sclereids.

Legumes

9.5–10.5 × 5.5–6 mm, short-hairy.

Seeds

1, reniform to ellipsoid.

dark red-brown, reniform-ellipsoid, 6–7 × 4–5 mm.

x

= 11.

Hoita

Hoita strobilina

Phenology Flowering Jun–Aug.
Habitat Woody slopes, usually on serpentine soils.
Elevation 100–900 m. (300–3000 ft.)
Distribution
from USDA
nw Mexico; California
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Species 3 (3 in the flora).

Species of Hoita are known from streamsides to moist banks and humid wooded slopes of coastal mountains and foothills of the Sierra Nevada.

J. W. Grimes (1990) recognized only three species in Hoita, reducing four of the 11 species recognized by Rydberg to synonymy under H. macrostachya, transferring one to Otholobium C. H. Stirton, and three to Rupertia. Hoita in the sense of Grimes is distinguished from other genera of Psoraleeae by the presence of the extra, internal wall in the fruit, and it is the only genus of the tribe in which the calyx does not enlarge or change shape through flowering and fruiting. Separation of Hoita from other Psoraleeae is further validated by a molecular phylogenetic study of the tribe (A. N. Egan and K. A. Crandall 2008).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hoita strobilina is known from Alameda, San Mateo, Santa Clara, and Santa Cruz counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems prostrate.
H. orbicularis
1. Stems erect.
→ 2
2. Stipules falcately lanceolate to caudate-elliptic, 7–16 mm, erect to reflexed.
H. strobilina
2. Stipules triangular to linear-triangular, 1.5–5 mm, erect.
H. macrostachya
Source FNA vol. 11. Author: Martin F. Wojciechowski. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae Fabaceae > subfam. Faboideae > Hoita
Sibling taxa
H. macrostachya, H. orbicularis
Subordinate taxa
H. macrostachya, H. orbicularis, H. strobilina
Synonyms Psoralea strobilina
Name authority Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 7. (1919) (Hooker & Arnott) Rydberg in N. L. Britton et al.: N. Amer. Fl. 24: 11. (1919)
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