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little brown jug

birthwort family, Dutchman's-pipe family, pipevine family

Habit Herbs or lianas [shrubs, rarely trees], deciduous or evergreen, often aromatic.
Rhizomes

internodes short, leaves crowded at rhizome apex (or internodes somewhat elongate, leaves scarcely crowded in var. arifolia and var. callifolia growing in wet places).

Leaves

blade commonly variegate, triangular-sagittate to subhastate, infrequently ovate-sagittate to deltate.

blade unlobed, margins entire.

Inflorescences

terminal or axillary, racemes or solitary flowers, rarely fan-shaped cymes.

Flowers

calyx tube narrowly to broadly urceolate-campanulate or ovoid, 12-30 × 6-12 mm, inner surface smooth;

lobes erect or spreading, 2-8 × 2-9 mm, adaxially puberulent;

stamen connective extending slightly beyond pollen sacs;

ovary ca. 1/3-inferior;

ovules ca. 6 per locule;

styles 2-cleft to stigma.

bisexual;

calyx enlarged, petaloid, usually tubular, [1-,] 3-, [6-, rarely 5-]merous, lobes valvate;

corolla usually reduced to scales or absent;

stamens 5, 6, or 12 [multiples of 3 or 5], free or adnate to styles and stigmas, forming gynostemium;

anthers extrorse;

pistil 1, 4-6-carpellate;

ovary inferior, partly inferior, or superior;

placentation axile (and ovaries 4-6-locular) or parietal;

ovules many per locule, anatropous.

Fruits

capsules [follicles], regularly to irregularly loculicidal, rarely indehiscent [septicidal].

Seeds

often flattened;

endosperm copious.

Wood

with broad medullary rays.

Hexastylis arifolia

Aristolochiaceae

Distribution
from FNA
AL; FL; GA; KY; LA; MS; NC; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
Primarily pantropical and subtropical
[BONAP county map]
Discussion

Varieties 3 (all in the flora).

Hexastylis arifolia is the most widespread species in the genus. Along the boundaries where the ranges of the varieties meet, intermediate specimens are occasionally found.

The Catawba tribe used Hexastylis arifolia (no varieties specified) medicinally for stomach pains, miscellaneous pains, heart trouble, and backaches; the Rappahannock, for treating whooping cough and asthma (D. E. Moerman 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 5, species ca. 600 (3 genera, 28 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyx lobes erect, 2–4 × 2–4 mm.
var. ruthii
1. Calyx lobes spreading, 2.5–8 × 3–9 mm.
→ 2
2. Calyx tube 13–18 × 6–10 mm.
var. arifolia
2. Calyx tube 20–25 × 10–12 mm.
var. callifolia
1. Calyx bilaterally symmetric, usually bent or curved; ovary inferior; stems erect, twining, or procumbent.
Aristolochia
1. Calyx radially symmetric, straight; ovary inferior, partly inferior, or superior; stems rhizomatous.
→ 2
2. Sepals distinct; anthers each with prominent terminal appendage; styles connate in column; ovary inferior.
Asarum
2. Sepals connate for most of length; anthers without terminal appendages; styles distinct (except sometimes at extreme base); ovary superior or ca. 1/3-inferior.
Hexastylis
Source FNA vol. 3. FNA vol. 3, p. 44. Authors: Kerry Barringer, Alan T. Whittemore.
Parent taxa Aristolochiaceae > Hexastylis
Sibling taxa
H. contracta, H. lewisii, H. minor, H. naniflora, H. rhombiformis, H. shuttleworthii, H. speciosa, H. virginica
Subordinate taxa
H. arifolia var. arifolia, H. arifolia var. callifolia, H. arifolia var. ruthii
Aristolochia, Asarum, Hexastylis
Synonyms Asarum arifolium
Name authority (Michaux) Small: Fl. S.E. U.S., 1132. (1903) Jussieu
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