Heuchera sanguinea
Heuchera richardsonii
coralbells
Richardson's alum-root
20–40 cm, short and sometimes long stipitate-glandular.
(7–)20–95 cm, densely long stipitate-glandular.
petiole long stipitate-glandular;
blade reniform to orbiculate, shallowly 5–7-lobed, 2–5.5 cm, base cordate, lobes rounded, margins dentate, apex acute to obtuse, surfaces long stipitate-glandular on veins abaxially, glabrous or sparsely long stipitate-glandular adaxially.
petiole densely or sparsely long or short stipitate-glandular;
blade broadly ovate or cordate, deeply 5–7-lobed, 2.5–10 cm, base cordate or nearly truncate, lobes rounded, margins dentate, apex acute, surfaces long stipitate-glandular abaxially, glabrous or long stipitate-glandular adaxially.
moderately dense to diffuse.
dense to diffuse.
hypanthium weakly bilaterally to radially symmetric, free 2.5–2.8 mm, dark pink to red, broadly campanulate or urceolate, 4–8 mm, short stipitate-glandular proximally, sparsely long stipitate-glandular distally;
sepals spreading, darker red-tipped, equal, 2–3 mm, apex oblong or rounded;
petals spreading, pink or cream, narrowly oblanceolate, unlobed, 1.2–1.8 mm (shorter than sepals), margins entire;
stamens included 1.5–3 mm;
styles included 1.5–3 mm, 1.5–2 mm, 0.1+ mm diam.
hypanthium strongly bilaterally symmetric, free 2–7 mm, green, campanulate, abruptly inflated distal to adnation to ovary, 5–14 mm, short stipitate-glandular;
sepals erect, green-tipped, equal, 1.3–4.2 mm, apex rounded (sinuses wider than petals);
petals erect, green or greenish white, rarely pink, narrowly spatulate, unlobed, 1.3–4 mm, margins finely dentate or coarsely fimbriate;
stamens 1.5 mm included to 4 mm exserted;
styles from 0.6 mm included to 0.3 mm exserted, 4–6 mm, to 0.1 mm diam.
ovoid, 4.5–6 mm, beaks divergent, not papillose.
ovoid, 7–14.5 mm, beaks divergent, not papillose.
dark brown, ellipsoid, 0.5–0.6 mm, bluntly spiny.
dark or very dark brown, ellipsoid, 0.6–0.9 mm.
= 14, 28.
Heuchera sanguinea
Heuchera richardsonii
Heuchera sanguinea occurs in the southwestern United States and northern Chihuahua, Mexico, and is widely cultivated throughout the cooler parts of North America.
Heuchera sanguinea varies considerably within populations in shape, lobing, and size of leaves, in amount of hairs, and in shape, form, and color of flowers. Rydberg recognized var. pulchra as characterized by denser inflorescences and longer hairs on its pedicels, inferior part of the ovary, and sepals. Mixed populations with individuals bearing longer and shorter hairs can be found throughout the range of the species, especially in Arizona. It seems best to recognize H. sanguinea as a variable species without infraspecific taxa.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Heuchera richardsonii intergrades with H. americana where their ranges overlap in Arkansas, Illinois, Indiana, Michigan, Missouri, and Oklahoma; the intergrading form is recognized here as H. americana var. hirsuticaulis.
Heuchera hispida (H. americana var. hispida here; see thereunder) was confused with H. richardsonii for almost a hundred years, until C. O. Rosendahl et al. (1933) pointed out that the plants from the Midwest then passing as H. hispida Pursh were distinct from Pursh’s species and were H. richardsonii.
The Blackfoot, Cree, Lakota, and Woodlands Indians used decoctions and infusions of the roots of Heuchera richardsonii for diarrhea and as an eyewash, and the Lakota applied a poultice of powdered roots to sores (D. E. Moerman 1998).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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