Heuchera sanguinea |
Heuchera grossulariifolia |
|
---|---|---|
coralbells |
gooseberry-leaf alumroot |
|
Habit | Herbs acaulescent; caudex branched. | Herbs acaulescent; caudex branched. |
Flowering stems | 20–40 cm, short and sometimes long stipitate-glandular. |
15–65 cm, glabrous or short stipitate-glandular. |
Leaves | petiole long stipitate-glandular; blade reniform to orbiculate, shallowly 5–7-lobed, 2–5.5 cm, base cordate, lobes rounded, margins dentate, apex acute to obtuse, surfaces long stipitate-glandular on veins abaxially, glabrous or sparsely long stipitate-glandular adaxially. |
petiole glabrous or short stipitate-glandular; blade cordate or orbiculate, shallowly 3–5-lobed, 1–7 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces glabrous or short stipitate-glandular. |
Inflorescences | moderately dense to diffuse. |
dense, interrupted. |
Flowers | hypanthium weakly bilaterally to radially symmetric, free 2.5–2.8 mm, dark pink to red, broadly campanulate or urceolate, 4–8 mm, short stipitate-glandular proximally, sparsely long stipitate-glandular distally; sepals spreading, darker red-tipped, equal, 2–3 mm, apex oblong or rounded; petals spreading, pink or cream, narrowly oblanceolate, unlobed, 1.2–1.8 mm (shorter than sepals), margins entire; stamens included 1.5–3 mm; styles included 1.5–3 mm, 1.5–2 mm, 0.1+ mm diam. |
hypanthium weakly bilaterally symmetric, free 1.2–1.9 mm, cream, broadly campanulate, 4–7 mm, short stipitate-glandular; sepals erect, often red-tipped, equal, 2 mm, apex rounded; petals erect, pink or white, spatulate to narrowly oblanceolate, (clawed), unlobed, 1–3 mm, margins entire; stamens included 1.5 mm; (filaments strongly incurved, shorter than and almost concealed by anthers); styles included to 1 mm, 0.5 mm, 0.1+ mm diam. |
Capsules | ovoid, 4.5–6 mm, beaks divergent, not papillose. |
ovoid, 5 mm, beaks divergent, not papillose. |
Seeds | dark brown, ellipsoid, 0.5–0.6 mm, bluntly spiny. |
dark brown, ellipsoid, 0.6 mm. |
2n | = 14, 28. |
|
Heuchera sanguinea |
Heuchera grossulariifolia |
|
Phenology | Flowering Mar–Oct. | Flowering May–Aug. |
Habitat | Moist, shaded rocks | Grassy hillsides, rocky canyon walls, alpine talus slopes |
Elevation | 1200-2600 m (3900-8500 ft) | 100-3400 m (300-11200 ft) |
Distribution |
AZ; NM; Mexico (Chihuahua)
|
ID; MT; NV; OR; WA
|
Discussion | Heuchera sanguinea occurs in the southwestern United States and northern Chihuahua, Mexico, and is widely cultivated throughout the cooler parts of North America. Heuchera sanguinea varies considerably within populations in shape, lobing, and size of leaves, in amount of hairs, and in shape, form, and color of flowers. Rydberg recognized var. pulchra as characterized by denser inflorescences and longer hairs on its pedicels, inferior part of the ovary, and sepals. Mixed populations with individuals bearing longer and shorter hairs can be found throughout the range of the species, especially in Arizona. It seems best to recognize H. sanguinea as a variable species without infraspecific taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Heuchera grossulariifolia includes both diploids and autotetraploids. K. A. Segraves and J. N. Thompson (1999) analyzed floral traits and flowering phenology in diploid and autotetraploid plants. Overall, plant size was greater in tetraploids than in diploids; flowers of tetraploids were larger (average hypanthium 6.5 mm) than those of diploids (average hypanthium 5.5 mm) and had a slightly different shape and phenology, but the diploids and tetraploids were not assigned taxonomic status in their study. Diploids and tetraploids were mixed in some populations, where characters intergraded (D. E. Soltis, pers. comm.). The autotetraploids have had two to seven independent origins from diploid progenitors, and do not represent a monophyletic lineage (Segraves and Thompson; Segraves et al. 1999). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 8, p. 100. | FNA vol. 8, p. 103. |
Parent taxa | Saxifragaceae > Heuchera | Saxifragaceae > Heuchera |
Sibling taxa | ||
Synonyms | H. sanguinea var. pulchra | H. cusickii, H. grossulariifolia var. tenuifolia, H. tenuifolia |
Name authority | Engelmann: in F. A. Wislizenus, Mem. Tour N. Mexico, 107. 1848 , | Rydberg: Mem. New York Bot. Gard. 1: 196. (1900) |
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