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island alumroot, jill-of-the-rocks

alumroot, American alum-root, common alum-root

Habit Herbs caulescent; caudex branched. Herbs acaulescent; caudex branched.
Flowering stems

9–30 cm, short or long stipitate-glandular, viscid.

leafy, 40–145 cm, glabrous or short stipitate-glandular.

Leaves

petiole (7–22 cm), densely long stipitate-glandular;

blade pentagonal to septagonal, deeply 5-lobed, 6.3–17 cm, base deeply cordate, lobes rounded, margins crenate, apex obtuse, surfaces long stipitate-glandular abaxially, glabrous adaxially except long stipitate-glandular near margins, viscid.

petiole glabrous or very short to long stipitate-glandular;

blade (often variegated adaxially), broadly ovate to cordate, shallowly 5–9-lobed, 3.5–11 cm, base cordate to nearly truncate, lobes rounded or ovate, margins dentate, apex acute or obtuse, surfaces abaxially glabrous or short stipitate-glandular, adaxially glabrous or short stipitate-glandular.

Inflorescences

dense.

diffuse.

Flowers

hypanthium radially symmetric, free 0.3–1.4 mm, white to pink, hemispheric, 2.5–4.2 mm, densely long stipitate-glandular;

sepals somewhat spreading, green- or red-tipped, equal, 0.7–2 mm, apex rounded or truncate, (medium to long stipitate-glandular);

petals spreading, white, oblong-lanceolate, (clawed), unlobed, 2–4 mm, margins entire;

stamens exserted 0.4–2.5 mm;

styles exserted to 2 mm, 1.5–4 mm, 0.1+ mm diam.

hypanthium weakly bilaterally symmetric, free 0.6–2 mm, green, urceolate or campanulate, abruptly inflated distal to adnation to ovary, 3–7.2 mm, very short stipitate-glandular;

sepals erect, green-tipped, equal, 1–2.4 mm, apex rounded;

petals erect, greenish, white, pink, or purple, narrowly spatulate, unlobed, 0.9–4 mm, margins entire or finely dentate or fimbriate;

stamens exserted 3–5 mm;

styles exserted 2.6–6.4 mm, 4–7 mm, to 0.1 mm diam.

Capsules

ovoid, 4–8 mm, beaks divergent, not papillose.

ovoid, 4–10.5 mm, beaks divergent, not papillose.

Seeds

black, broadly fusiform, 0.4 mm.

dark brown, ellipsoid, 0.6–0.9 mm.

2n

= 14.

Heuchera maxima

Heuchera americana

Phenology Flowering Mar–May.
Habitat Exposed cliffs
Elevation 0-500 m (0-1600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; GA; IL; IN; KY; LA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; VA; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Heuchera maxima is found on the northern Channel Islands (Anacapa, Santa Cruz, Santa Rosa), Santa Barbara County.

Heuchera maxima is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Heuchera americana is highly polymorphic and distributed over a large part of the eastern United States and Canada. Its variability is largely the result of its wide geographic range, the sporadic, semi-isolated distribution of populations, and interaction of differential adaptation and genetic drift made possible by its distribution pattern.

Heuchera americana intergrades with both H. pubescens and H. richardsonii where they overlap; the intergrading form with H. pubescens is H. americana var. hispida, and with H. richardsonii it is H. americana var. hirsuticaulis. A breeding study between H. americana, H. pubescens, H. richardsonii, and other species demonstrated ease of artificial hybridization and fertility of offspring among H. americana, H. pubescens, and H. richardsonii (E. F. Wells 1979).

Individuals of the three varieties do not form intermixed populations; populations tend to be geographically isolated from one another and to be relatively uniform, displaying somewhat narrow character variation within a population.

The Cherokee Indians took Heuchera americana for dysentery and used the powdered root for malignant ulcers, bad sores, bowel complaints, piles, female problems, and sore mouth. The Chickasaw Indians used the root as an astringent and tonic (D. E. Moerman 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petioles densely long stipitate-glandular.
var. hirsuticaulis
1. Petioles glabrous or short to sparsely long stipitate-glandular
→ 2
2. Petioles glabrous or very short to sparsely long stipitate-glandular; hypanthia free 0.6-1.5 mm; petals greenish, white, or pink, narrower than sepals, margins entire or finely dentate.
var. americana
2. Petioles glabrous or very short stipitate-glandular; hypanthia free 1.5-2 mm; petals purple or pink, wider than sepals, margins fimbriate.
var. hispida
Source FNA vol. 8, p. 93. FNA vol. 8, p. 94.
Parent taxa Saxifragaceae > Heuchera Saxifragaceae > Heuchera
Sibling taxa
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. grossulariifolia, H. hallii, H. hirsutissima, H. longiflora, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
H. abramsii, H. alba, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. grossulariifolia, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
Subordinate taxa
H. americana var. americana, H. americana var. hirsuticaulis, H. americana var. hispida
Name authority Greene: Bull. Calif. Acad. Sci. 2: 149. 1886 , Linnaeus: Sp. Pl. 1: 226. 1753 ,
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