Heuchera hirsutissima
Heuchera americana
shaggyhair alum-root
alumroot, American alum-root, common alum-root
10–40 cm, short stipitate-glandular and sparsely long stipitate-glandular.
leafy, 40–145 cm, glabrous or short stipitate-glandular.
petiole long stipitate-glandular;
blade orbiculate to reniform, shallowly 5-lobed, 1–1.5 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces long stipitate-glandular.
petiole glabrous or very short to long stipitate-glandular;
blade (often variegated adaxially), broadly ovate to cordate, shallowly 5–9-lobed, 3.5–11 cm, base cordate to nearly truncate, lobes rounded or ovate, margins dentate, apex acute or obtuse, surfaces abaxially glabrous or short stipitate-glandular, adaxially glabrous or short stipitate-glandular.
diffuse.
hypanthium strongly bilaterally symmetric, free 0.8 mm on abaxial side, 1.8 mm on adaxial side, pink, narrowly cylindric, 5–6.5 mm, short stipitate-glandular proximally, densely long stipitate-glandular distally;
sepals spreading, green- and red-tipped, unequal, 0.5–1.5 mm on abaxial side of hypanthium, to 0.5 mm on adaxial side, apex rounded;
petals spreading, white, oblanceolate, (clawed), unlobed, 2–2.5 mm (equaling or longer than sepals, rarely shorter), margins entire;
stamens exserted to 0.5 mm;
styles included to 0.5 mm, 1.5 mm, 0.1+ mm diam.
hypanthium weakly bilaterally symmetric, free 0.6–2 mm, green, urceolate or campanulate, abruptly inflated distal to adnation to ovary, 3–7.2 mm, very short stipitate-glandular;
sepals erect, green-tipped, equal, 1–2.4 mm, apex rounded;
petals erect, greenish, white, pink, or purple, narrowly spatulate, unlobed, 0.9–4 mm, margins entire or finely dentate or fimbriate;
stamens exserted 3–5 mm;
styles exserted 2.6–6.4 mm, 4–7 mm, to 0.1 mm diam.
ovoid, 3 mm, beaks divergent, not papillose.
ovoid, 4–10.5 mm, beaks divergent, not papillose.
dark brown, ellipsoid, ca. 0.7 mm.
dark brown, ellipsoid, 0.6–0.9 mm.
dense.
= 14.
Heuchera hirsutissima
Heuchera americana
Of conservation concern.
Heuchera hirsutissima is known from Mount San Jacinto and the San Jacinto Mountains of Riverside County and possibly the San Bernardino Mountains of San Bernardino County. It resembles H. caespitosa and H. elegans, except for its densely villous hypanthium and minute adaxial sepals.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 3 (3 in the flora).
Heuchera americana is highly polymorphic and distributed over a large part of the eastern United States and Canada. Its variability is largely the result of its wide geographic range, the sporadic, semi-isolated distribution of populations, and interaction of differential adaptation and genetic drift made possible by its distribution pattern.
Heuchera americana intergrades with both H. pubescens and H. richardsonii where they overlap; the intergrading form with H. pubescens is H. americana var. hispida, and with H. richardsonii it is H. americana var. hirsuticaulis. A breeding study between H. americana, H. pubescens, H. richardsonii, and other species demonstrated ease of artificial hybridization and fertility of offspring among H. americana, H. pubescens, and H. richardsonii (E. F. Wells 1979).
Individuals of the three varieties do not form intermixed populations; populations tend to be geographically isolated from one another and to be relatively uniform, displaying somewhat narrow character variation within a population.
The Cherokee Indians took Heuchera americana for dysentery and used the powdered root for malignant ulcers, bad sores, bowel complaints, piles, female problems, and sore mouth. The Chickasaw Indians used the root as an astringent and tonic (D. E. Moerman 1998).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Petioles densely long stipitate-glandular. | var. hirsuticaulis |
1. Petioles glabrous or short to sparsely long stipitate-glandular | → 2 |
2. Petioles glabrous or very short to sparsely long stipitate-glandular; hypanthia free 0.6-1.5 mm; petals greenish, white, or pink, narrower than sepals, margins entire or finely dentate. | var. americana |
2. Petioles glabrous or very short stipitate-glandular; hypanthia free 1.5-2 mm; petals purple or pink, wider than sepals, margins fimbriate. | var. hispida |
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