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gooseberry-leaf alumroot

hairy alum-root, maple-leaf alum-root

Habit Herbs acaulescent; caudex branched. Herbs caulescent; caudex branched.
Flowering stems

15–65 cm, glabrous or short stipitate-glandular.

10–85 cm, long stipitate-glandular.

Leaves

petiole glabrous or short stipitate-glandular;

blade cordate or orbiculate, shallowly 3–5-lobed, 1–7 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces glabrous or short stipitate-glandular.

petiole long stipitate-glandular;

blade rounded-cordate, orbiculate, or oblong, often ± polygonal, deeply (5–)7–9+-lobed, 3–26 cm, base cordate to nearly truncate, lobes ovate to triangular, margins dentate to serrate, apex obtuse to acute, surfaces glabrous or long stipitate-glandular.

Inflorescences

dense, interrupted.

dense to diffuse.

Flowers

hypanthium weakly bilaterally symmetric, free 1.2–1.9 mm, cream, broadly campanulate, 4–7 mm, short stipitate-glandular;

sepals erect, often red-tipped, equal, 2 mm, apex rounded;

petals erect, pink or white, spatulate to narrowly oblanceolate, (clawed), unlobed, 1–3 mm, margins entire;

stamens included 1.5 mm; (filaments strongly incurved, shorter than and almost concealed by anthers);

styles included to 1 mm, 0.5 mm, 0.1+ mm diam.

hypanthium radially symmetric, free 0.2–0.5 mm, white or pink, obconic or subglobose, 1.5–3.6 mm, usually long stipitate-glandular, sometimes sparsely long-stipitate or glabrous;

sepals erect, green-tipped, equal, 0.5–1.7 mm, apex rounded;

petals often coiled, sometimes reflexed, white, linear, lanceolate, or oblanceolate, unlobed, 1–3 mm, margins entire;

stamens exserted 2–2.5 mm;

styles exserted 1.5–2.5 mm, 2–4 mm, to 0.1 mm diam.

Capsules

ovoid, 5 mm, beaks divergent, not papillose.

ovoid, 3–6.4 mm, beaks divergent, not papillose.

Seeds

dark brown, ellipsoid, 0.6 mm.

dark brown, fusiform, 0.5–0.9 mm.

2n

= 14, 28.

Heuchera grossulariifolia

Heuchera villosa

Phenology Flowering May–Aug.
Habitat Grassy hillsides, rocky canyon walls, alpine talus slopes
Elevation 100-3400 m (300-11200 ft)
Distribution
from FNA
ID; MT; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; GA; IN; KY; MO; MS; NC; OH; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
Discussion

Heuchera grossulariifolia includes both diploids and autotetraploids. K. A. Segraves and J. N. Thompson (1999) analyzed floral traits and flowering phenology in diploid and autotetraploid plants. Overall, plant size was greater in tetraploids than in diploids; flowers of tetraploids were larger (average hypanthium 6.5 mm) than those of diploids (average hypanthium 5.5 mm) and had a slightly different shape and phenology, but the diploids and tetraploids were not assigned taxonomic status in their study. Diploids and tetraploids were mixed in some populations, where characters intergraded (D. E. Soltis, pers. comm.). The autotetraploids have had two to seven independent origins from diploid progenitors, and do not represent a monophyletic lineage (Segraves and Thompson; Segraves et al. 1999).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences dense to diffuse, internodes in floral branchlets 0.7-3 mm; pedicels 0.9-3 mm; hypanthia obconic or subglobose, long stipitate-glandular; petals 0.1-0.2 mm wide; e of the Prairies.
var. villosa
1. Inflorescences dense, internodes in floral branchlets 0.3-1 mm; pedicels 0.2-0.9 mm; hypanthia subglobose, glabrous or sparsely long stipitate-glandular; petals 0.2-0.4 mm wide; Arkansas.
var. arkansana
Source FNA vol. 8, p. 103. FNA vol. 8, p. 89.
Parent taxa Saxifragaceae > Heuchera Saxifragaceae > Heuchera
Sibling taxa
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. grossulariifolia, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. wootonii
Subordinate taxa
H. villosa var. arkansana, H. villosa var. villosa
Synonyms H. cusickii, H. grossulariifolia var. tenuifolia, H. tenuifolia
Name authority Rydberg: Mem. New York Bot. Gard. 1: 196. (1900) Michaux: Fl. Bor.-Amer. 1: 172. 1803 ,
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