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gooseberry-leaf alumroot

long-flower alum-root

Habit Herbs acaulescent; caudex branched. Herbs acaulescent; caudex branched.
Flowering stems

15–65 cm, glabrous or short stipitate-glandular.

often leafy, 30–95 cm, glabrous or short stipitate-glandular.

Leaves

petiole glabrous or short stipitate-glandular;

blade cordate or orbiculate, shallowly 3–5-lobed, 1–7 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces glabrous or short stipitate-glandular.

petiole glabrous or short stipitate-glandular;

blade (often variegated adaxially), broadly ovate to cordate, shallowly 5-lobed, 3–12 cm, base cordate or nearly truncate, lobes rounded to widely ovate, terminal lobes often elongated, margins dentate, apex acute or obtuse, surfaces glabrous or short stipitate-glandular, at least on veins.

Inflorescences

dense, interrupted.

diffuse.

Flowers

hypanthium weakly bilaterally symmetric, free 1.2–1.9 mm, cream, broadly campanulate, 4–7 mm, short stipitate-glandular;

sepals erect, often red-tipped, equal, 2 mm, apex rounded;

petals erect, pink or white, spatulate to narrowly oblanceolate, (clawed), unlobed, 1–3 mm, margins entire;

stamens included 1.5 mm; (filaments strongly incurved, shorter than and almost concealed by anthers);

styles included to 1 mm, 0.5 mm, 0.1+ mm diam.

hypanthium strongly bilaterally symmetric, free 2.2–6.2 mm, green, gibbous-tubular, abruptly inflated distal to adnation to ovary, 6.6–12.6 mm, short stipitate-glandular;

sepals inflexed (closing mouth of flower), darker green-tipped, equal, 2–3.7 mm, apex rounded;

petals inflexed (closing mouth of flower), white, pink, or purple, spatulate, unlobed, 1.8–5.5 mm, margins often fimbriate;

stamens 2.4 mm included to 0.7 mm exserted;

styles included 1.3–5.3 mm, 1.5–2.5 mm, to 0.1 mm diam.

Capsules

ovoid, 5 mm, beaks divergent, not papillose.

ovoid, 5–13 mm, beaks divergent, not papillose.

Seeds

dark brown, ellipsoid, 0.6 mm.

dark brown, ellipsoid, 0.5–0.9 mm.

2n

= 14, 28.

= 14.

Heuchera grossulariifolia

Heuchera longiflora

Phenology Flowering May–Aug. Flowering May–Jun.
Habitat Grassy hillsides, rocky canyon walls, alpine talus slopes Rich, shaded woods and roadcuts over limestone substrates and outcroppings
Elevation 100-3400 m (300-11200 ft) 100-500 m (300-1600 ft)
Distribution
from FNA
ID; MT; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; KY; NC; TN; VA; WV
[BONAP county map]
Discussion

Heuchera grossulariifolia includes both diploids and autotetraploids. K. A. Segraves and J. N. Thompson (1999) analyzed floral traits and flowering phenology in diploid and autotetraploid plants. Overall, plant size was greater in tetraploids than in diploids; flowers of tetraploids were larger (average hypanthium 6.5 mm) than those of diploids (average hypanthium 5.5 mm) and had a slightly different shape and phenology, but the diploids and tetraploids were not assigned taxonomic status in their study. Diploids and tetraploids were mixed in some populations, where characters intergraded (D. E. Soltis, pers. comm.). The autotetraploids have had two to seven independent origins from diploid progenitors, and do not represent a monophyletic lineage (Segraves and Thompson; Segraves et al. 1999).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Heuchera longiflora is restricted to limestone outcroppings and is rare in all the states where it is found.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 103. FNA vol. 8, p. 97.
Parent taxa Saxifragaceae > Heuchera Saxifragaceae > Heuchera
Sibling taxa
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. grossulariifolia, H. hallii, H. hirsutissima, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
Synonyms H. cusickii, H. grossulariifolia var. tenuifolia, H. tenuifolia H. aceroides, H. longiflora var. aceroides, H. scabra
Name authority Rydberg: Mem. New York Bot. Gard. 1: 196. (1900) Rydberg: in N. L. Britton, Man. Fl. N. States, 482. (1901)
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