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gooseberry-leaf alumroot

urn-flower alum-root

Habit Herbs acaulescent; caudex branched. Herbs subcaulescent; caudex branched.
Flowering stems

15–65 cm, glabrous or short stipitate-glandular.

10–40 cm, sparsely medium stipitate-glandular.

Leaves

petiole glabrous or short stipitate-glandular;

blade cordate or orbiculate, shallowly 3–5-lobed, 1–7 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces glabrous or short stipitate-glandular.

petiole long stipitate-glandular;

blade reniform or orbiculate, shallowly 5-lobed, 1.5–3.5 cm, base cordate or truncate, lobes rounded, margins dentate, apex obtuse, surfaces short or long stipitate-glandular.

Inflorescences

dense, interrupted.

Flowers

hypanthium weakly bilaterally symmetric, free 1.2–1.9 mm, cream, broadly campanulate, 4–7 mm, short stipitate-glandular;

sepals erect, often red-tipped, equal, 2 mm, apex rounded;

petals erect, pink or white, spatulate to narrowly oblanceolate, (clawed), unlobed, 1–3 mm, margins entire;

stamens included 1.5 mm; (filaments strongly incurved, shorter than and almost concealed by anthers);

styles included to 1 mm, 0.5 mm, 0.1+ mm diam.

hypanthium strongly bilaterally symmetric, free 3–3.5 mm on adaxial side, pink to purplish, narrowly cylindric or cylindric-urceolate, 4–7 mm, short stipitate-glandular proximally, densely long stipitate-glandular distally;

sepals spreading, green-tipped, unequal, 2 mm on adaxial side of hypanthium, apex obtuse or rounded;

petals spreading, white, broadly oblanceolate, (clawed), unlobed, 3–4 mm (longer than sepals), margins entire;

stamens included 0.5 mm to exserted 0.5 mm;

styles included to 2 mm, 1.5–2 mm, 0.1+ mm diam.

Capsules

ovoid, 5 mm, beaks divergent, not papillose.

urceolate, 4–6 mm, beaks divergent, not papillose.

Seeds

dark brown, ellipsoid, 0.6 mm.

dark brown, ellipsoid, 0.7 mm.

Inflorescenses

dense.

2n

= 14, 28.

Heuchera grossulariifolia

Heuchera elegans

Phenology Flowering May–Aug. Flowering May–Jun.
Habitat Grassy hillsides, rocky canyon walls, alpine talus slopes Rocky sites in yellow pine and red fir forests
Elevation 100-3400 m (300-11200 ft) 1500-2600 m (4900-8500 ft)
Distribution
from FNA
ID; MT; NV; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
Discussion

Heuchera grossulariifolia includes both diploids and autotetraploids. K. A. Segraves and J. N. Thompson (1999) analyzed floral traits and flowering phenology in diploid and autotetraploid plants. Overall, plant size was greater in tetraploids than in diploids; flowers of tetraploids were larger (average hypanthium 6.5 mm) than those of diploids (average hypanthium 5.5 mm) and had a slightly different shape and phenology, but the diploids and tetraploids were not assigned taxonomic status in their study. Diploids and tetraploids were mixed in some populations, where characters intergraded (D. E. Soltis, pers. comm.). The autotetraploids have had two to seven independent origins from diploid progenitors, and do not represent a monophyletic lineage (Segraves and Thompson; Segraves et al. 1999).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Heuchera elegans occurs in the mountains of southern California, chiefly in the San Gabriel Range in Los Angeles and San Bernardino counties and vicinity. It resembles H. caespitosa. This group is in need of phylogenetic study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 103. FNA vol. 8, p. 100.
Parent taxa Saxifragaceae > Heuchera Saxifragaceae > Heuchera
Sibling taxa
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. elegans, H. glabra, H. glomerulata, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
H. abramsii, H. alba, H. americana, H. bracteata, H. brevistaminea, H. caespitosa, H. caroliniana, H. chlorantha, H. cylindrica, H. eastwoodiae, H. glabra, H. glomerulata, H. grossulariifolia, H. hallii, H. hirsutissima, H. longiflora, H. maxima, H. merriamii, H. micrantha, H. novamexicana, H. parishii, H. parviflora, H. parvifolia, H. pilosissima, H. pubescens, H. pulchella, H. richardsonii, H. rubescens, H. sanguinea, H. villosa, H. wootonii
Synonyms H. cusickii, H. grossulariifolia var. tenuifolia, H. tenuifolia
Name authority Rydberg: Mem. New York Bot. Gard. 1: 196. (1900) Abrams: Bull. S. Calif. Acad. Sci. 1: 67. 1902 ,
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