Heuchera grossulariifolia
Heuchera caespitosa
gooseberry-leaf alumroot
tuft alumroot, urn-flower alum-root
15–65 cm, glabrous or short stipitate-glandular.
10–40 cm, long stipitate-glandular.
petiole glabrous or short stipitate-glandular;
blade cordate or orbiculate, shallowly 3–5-lobed, 1–7 cm, base cordate, lobes rounded, margins dentate, apex obtuse, surfaces glabrous or short stipitate-glandular.
petiole long stipitate-glandular;
blade reniform or orbiculate, shallowly 5-lobed, 1–4 cm, base cordate or truncate, lobes rounded, margins dentate, apex obtuse, surfaces short or long stipitate-glandular.
dense, interrupted.
hypanthium weakly bilaterally symmetric, free 1.2–1.9 mm, cream, broadly campanulate, 4–7 mm, short stipitate-glandular;
sepals erect, often red-tipped, equal, 2 mm, apex rounded;
petals erect, pink or white, spatulate to narrowly oblanceolate, (clawed), unlobed, 1–3 mm, margins entire;
stamens included 1.5 mm; (filaments strongly incurved, shorter than and almost concealed by anthers);
styles included to 1 mm, 0.5 mm, 0.1+ mm diam.
hypanthium strongly bilaterally symmetric, free 2–2.8 mm on adaxial side, pink to purplish, narrowly campanulate, widening at mouth, 4–7 mm, short stipitate-glandular proximally, long stipitate-glandular distally;
sepals spreading, green-tipped, equal, 1–1.2 mm, apex rounded;
petals spreading, white, oblanceolate, (clawed), unlobed, 2–2.5 mm (longer than sepals), margins entire;
stamens included 0.5 mm to exserted 0.5 mm;
styles to 0.5 mm included, 1.5 mm, 0.1+ mm diam.
ovoid, 5 mm, beaks divergent, not papillose.
ovoid, 4 mm, beaks divergent, not papillose.
dark brown, ellipsoid, 0.6 mm.
dark brown, ellipsoid, 0.7 mm.
moderately dense.
= 14, 28.
Heuchera grossulariifolia
Heuchera caespitosa
Heuchera grossulariifolia includes both diploids and autotetraploids. K. A. Segraves and J. N. Thompson (1999) analyzed floral traits and flowering phenology in diploid and autotetraploid plants. Overall, plant size was greater in tetraploids than in diploids; flowers of tetraploids were larger (average hypanthium 6.5 mm) than those of diploids (average hypanthium 5.5 mm) and had a slightly different shape and phenology, but the diploids and tetraploids were not assigned taxonomic status in their study. Diploids and tetraploids were mixed in some populations, where characters intergraded (D. E. Soltis, pers. comm.). The autotetraploids have had two to seven independent origins from diploid progenitors, and do not represent a monophyletic lineage (Segraves and Thompson; Segraves et al. 1999).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Of conservation concern.
Heuchera caespitosa occurs in the western Transverse Ranges in Kern, San Bernardino, Tulare, and Ventura counties, and in the Outer South Coast Ranges, southern Sierra Nevada foothills, and southern Sierra Nevada. It is very similar to H. abramsii, H. brevistaminea, H. hirsutissima, and H. pulchella; the group needs thorough phylogenetic analysis.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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WildflowerSearch
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