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golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster

camphor false golden-aster, camphorweed, golden aster, unknown

Habit Perennials, (5–)16–40(–70) cm; taprooted. Annuals or biennials, 10–200 cm (aromatic); taprooted (rarely perennating in south from proximal stem nodes).
Stems

1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular.

1–4+, procumbent to erect (sometimes reddish brown, proximal to distal branches well developed in larger plants), sparsely to densely hispido-strigose, sparsely to densely stipitate-glandular.

Leaves

generally not crowded;

proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular;

distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular.

basal and proximal rarely persisting to flowering, sometimes present and withered, brown to black;

basal to mid cauline petiolate (petioles 10–40 mm, bases auriculate-clasping), blades ovate to elliptic or lanceolate, 10–70 × 6–55 mm, bases cuneate to attenuate, margins flat or undulate, coarsely serrate or entire, basally long-ciliate, apices acute, faces moderately hispido-scabrous;

distal sessile, blades ovate to lanceolate, 10–90 × 2–20 mm, reduced distally, bases often becoming cordate distally, subclasping or not clasping, margins entire, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular.

Peduncles

(4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular;

bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads.

2–35 mm, sparsely to densely hispido-strigose, moderately to densely stipitate-glandular;

bracts 0–4, proximal leaflike, ovate to lanceolate, reduced distally and becoming linear, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular.

Involucres

narrowly cylindric to campanulate, (5–)6–9.5(–13) mm.

hemispheric to campanulate, 4–8(–10) mm.

Ray florets

(5–)10–20(–38);

laminae (3.5–)6.5–11(–20) × 1–2(–3) mm.

15–35;

laminae 3–7(–9) × 1–2 mm.

Disc florets

(10–)20–50(–85);

corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm).

25–60;

corollas weakly ampliate, 2–9 mm, proximal throats glabrous to sparsely short-strigose, lobes 0.5–0.7 mm, glabrous.

Phyllaries

in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular.

in 4–6 series, lanceolate, sometimes strongly unequal, margins scarious and distally strigoso-ciliate, faces sparsely to densely stipitate-glandular, with 0–28 coarse, scabro-strigose hairs distally.

Heads

1–16(–42), in usually open, corymbiform, rarely paniculiform arrays.

3–180+ in corymbo-paniculiform arrays, sometimes becoming profusely branched (loosely paniculiform or broadly corymbiform in larger plants), branches ascending to spreading.

Cypselae

monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose;

pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas).

dimorphic, obconic, ribs 2–3, (ray) triangular in cross section, 1.5–2.5 mm, faces glabrous to slightly strigose, (disc) laterally compressed, 2–4 mm, faces moderately to densely strigose;

pappi 0 (ray) or (disc) tan to rust, outer of linear to triangular scales 0.25–0.6 mm, inner of 25–45 bristles 4–9 mm, longest weakly clavate.

2n

= 18, 36.

= 18.

Heterotheca villosa

Heterotheca subaxillaris

Distribution
from FNA
AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OK; PA; SC; TN; TX; UT; VA; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 9 (9 in the flora).

Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Heterotheca subaxillaris is a very weedy, morphologically variable species that has been treated as three species (B. L. Wagenknecht 1960; V. L. Harms 1965, 1970; J. C. Semple 1996) or as a single polymorphic species (G. L. Nesom 1990e). Harms (1965) acknowledged the difficulty in finding consistent characters to separate plants on a geographic basis, as did Nesom (1990e). Harms (1970) noted that H. psammophila and H. latifolia are probably conspecific. It seems best to treat the complex as a single polymorphic species. In historic times, human-assisted dispersal of possibly once distinct races has likely resulted in blurring of such distinctions with one exception. Plants of the outer coastal plain from Texas to North Carolina usually have a cluster of coarse hairs near the tip of phyllaries, at least in summer and fall-blooming plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems sparsely to densely hispido-strigose, sparsely to densely stipitate-glandular; distal cauline leaves sparsely to densely hispido-strigose (hairs 2–100/mm²), sparsely to densely stipitate-glandular (glands 2–60/mm²)
→ 2
1. Stems moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular; distal leaf faces moderately to densely strigose (hairs 25–200/mm²), eglandular or sparsely stipitate-glandular (rarely more densely so)
→ 5
2. Distal cauline leaf blades ovate to oblanceolate or lanceolate to triangular-lanceolate, rarely oblong, glands sometimes more numerous than hairs/mm²; arrays corymbiform
→ 3
2. Distal cauline leaf blades linear, lanceolate, narrowly to broadly oblong, or narrowly ovate, glands less numerous than hairs/mm²; arrays congested, corymbiform to subumbelliform; w Great Plains, foothills of Front Range, White Mountains (New Mexico)
→ 4
3. Distal cauline leaf blades usually narrowly to broadly oblanceolate, sometimes oblong, sometimes ovate in mountains, margins flat to remotely undulate, apices obtuse to acute, indument sparse to moderate; arrays compact, branches usually not very long; phyllaries usually moderately strigose (hairs to 1 mm), often stipitate-glandular
var. minor
3. Distal cauline leaf blades narrowly lanceolate to triangular-lanceolate, apices sharply acute, margins flat; arrays lax, branches often very long; phyllaries sparsely short-strigose, hairs usually less than 0.3 mm, sparsely to moderately stipitate-glandular; rock outcrops, margins of arroyos, rocky banks, Arizona, Idaho, Nevada, Utah, disjunct in Little San Bernardino Mountains, California
var. scabra
4. Distal leaf blades often perpendicular to stems, linear to broadly oblong, sometimes small (less than 10 mm), faces moderately hispido-strigose (hairs 9–56/mm²), moderately to densely stipitate-glandular (glands 9–36/mm²), glands sometimes more than hairs; ray laminae (3.5–)6.9–11(–12) mm; lower elevations in mountains and foothills, Colorado and New Mexico, Great Plains, Colorado, w Nebraska, New Mexico, se Wyoming
var. nana
4. Distal leaf blades (perpendicular proximally, ascending distally) lanceolate to ovate-oblong, faces moderately to densely strigose (hairs usually 42–93/mm²), sparsely to moderately stipitate-glandular (glands usually 4–37/mm²); ray laminae (8.5–)9–12(–14) mm; mid elevations of Sierra Blanca, New Mexico, and mountains to s, w
var. sierrablancensis
5. Stems often abundantly long-hirsute; distal cauline leaf blades narrowly to broadly oblong; bracts subtending heads often surpassing them
→ 6
5. Stems sparsely to moderately long-hirsute; distal cauline leaf blades narrowly to broadly oblanceolate; bracts subtending heads small or none
→ 7
6. Leaf faces moderately strigose (hairs usually not obscuring faces); involucres broadly campanulate; rays showy, (12–)17–30 (averaging 21), laminae (8.5–)9.9–14.5 mm (averaging 11.5 mm); ne Great Plains
var. ballardii
6. Leaf faces moderately to densely hispido-strigose (hairs sometimes obscuring faces); involucres narrowly campanulate; rays less showy, (7–)13–22 (averaging 17), laminae 8–11.4(–12.4) mm (averaging 10 mm); mountains and w prairies
var. foliosa
7. Stems moderately to densely strigose, sparsely to abundantly long-hirsute; distal cauline leaf faces moderately strigose (20–60 hairs/mm²); Great Plains, scattered in hills and mountains
var. villosa
7. Stems sparsely long-hirsute; distal cauline leaf faces densely strigose (60–270 hairs/mm²)
→ 8
8. Stems 5–22 cm, sparsely long-hirsute; distal leaf faces densely strigose (84–140 hairs/mm²), sparsely to moderately stipitate-glandular; geyser basins, Yellowstone National Park, gravelly stream banks, e Teton National Park, rare further s in Wyoming
var. depressa
8. Stems (12–)30–50(–75) cm, usually sparsely, rarely abundantly long-hirsute; distal leaf faces densely strigose (60–270 hairs/mm²), eglandular or sparsely stipitate-glandular; grasslands, wooded hillsides, mountains, Arizona, Colorado, New Mexico, Utah
var. pedunculata
1. Outer and mid phyllaries with apical tuft of 6–28 coarse, scabro-strigose hairs; coastal plain, Texas to Delaware
subsp. subaxillaris
1. Outer and mid phyllaries without distinct apical tuft of hairs (sometimes distally with 1–2 coarse, scabro-strigose hairs); throughout s United States
subsp. latifolia
Source FNA vol. 20, p. 2. FNA vol. 20, p. 234.
Parent taxa Asteraceae > tribe Astereae > Heterotheca Asteraceae > tribe Astereae > Heterotheca
Sibling taxa
H. camporum, H. canescens, H. fulcrata, H. grandiflora, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. stenophylla, H. subaxillaris, H. viscida, H. zionensis
H. camporum, H. canescens, H. fulcrata, H. grandiflora, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. stenophylla, H. villosa, H. viscida, H. zionensis
Subordinate taxa
H. villosa var. ballardii, H. villosa var. depressa, H. villosa var. foliosa, H. villosa var. minor, H. villosa var. nana, H. villosa var. pedunculata, H. villosa var. scabra, H. villosa var. sierrablancensis, H. villosa var. villosa
H. subaxillaris subsp. latifolia, H. subaxillaris subsp. subaxillaris
Synonyms Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus Inula subaxillaris
Name authority (Pursh) Shinners: Field & Lab. 19: 71. (1951) (Lamarck) Britton & Rusby: Trans. New York Acad. Sci. 7: 10. (1887)
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