Heterotheca villosa |
Heterotheca subaxillaris |
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golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster |
camphor false golden-aster, camphorweed, golden aster, unknown |
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Habit | Perennials, (5–)16–40(–70) cm; taprooted. | Annuals or biennials, 10–200 cm (aromatic); taprooted (rarely perennating in south from proximal stem nodes). | ||||||||||||||||||||||||||||||||||||
Stems | 1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular. |
1–4+, procumbent to erect (sometimes reddish brown, proximal to distal branches well developed in larger plants), sparsely to densely hispido-strigose, sparsely to densely stipitate-glandular. |
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Leaves | generally not crowded; proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular. |
basal and proximal rarely persisting to flowering, sometimes present and withered, brown to black; basal to mid cauline petiolate (petioles 10–40 mm, bases auriculate-clasping), blades ovate to elliptic or lanceolate, 10–70 × 6–55 mm, bases cuneate to attenuate, margins flat or undulate, coarsely serrate or entire, basally long-ciliate, apices acute, faces moderately hispido-scabrous; distal sessile, blades ovate to lanceolate, 10–90 × 2–20 mm, reduced distally, bases often becoming cordate distally, subclasping or not clasping, margins entire, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular. |
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Peduncles | (4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular; bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads. |
2–35 mm, sparsely to densely hispido-strigose, moderately to densely stipitate-glandular; bracts 0–4, proximal leaflike, ovate to lanceolate, reduced distally and becoming linear, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular. |
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Involucres | narrowly cylindric to campanulate, (5–)6–9.5(–13) mm. |
hemispheric to campanulate, 4–8(–10) mm. |
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Ray florets | (5–)10–20(–38); laminae (3.5–)6.5–11(–20) × 1–2(–3) mm. |
15–35; laminae 3–7(–9) × 1–2 mm. |
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Disc florets | (10–)20–50(–85); corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm). |
25–60; corollas weakly ampliate, 2–9 mm, proximal throats glabrous to sparsely short-strigose, lobes 0.5–0.7 mm, glabrous. |
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Phyllaries | in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular. |
in 4–6 series, lanceolate, sometimes strongly unequal, margins scarious and distally strigoso-ciliate, faces sparsely to densely stipitate-glandular, with 0–28 coarse, scabro-strigose hairs distally. |
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Heads | 1–16(–42), in usually open, corymbiform, rarely paniculiform arrays. |
3–180+ in corymbo-paniculiform arrays, sometimes becoming profusely branched (loosely paniculiform or broadly corymbiform in larger plants), branches ascending to spreading. |
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Cypselae | monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas). |
dimorphic, obconic, ribs 2–3, (ray) triangular in cross section, 1.5–2.5 mm, faces glabrous to slightly strigose, (disc) laterally compressed, 2–4 mm, faces moderately to densely strigose; pappi 0 (ray) or (disc) tan to rust, outer of linear to triangular scales 0.25–0.6 mm, inner of 25–45 bristles 4–9 mm, longest weakly clavate. |
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2n | = 18, 36. |
= 18. |
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Heterotheca villosa |
Heterotheca subaxillaris |
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Distribution |
AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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AL; AR; AZ; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OK; PA; SC; TN; TX; UT; VA; Mexico
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Discussion | Varieties 9 (9 in the flora). Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Heterotheca subaxillaris is a very weedy, morphologically variable species that has been treated as three species (B. L. Wagenknecht 1960; V. L. Harms 1965, 1970; J. C. Semple 1996) or as a single polymorphic species (G. L. Nesom 1990e). Harms (1965) acknowledged the difficulty in finding consistent characters to separate plants on a geographic basis, as did Nesom (1990e). Harms (1970) noted that H. psammophila and H. latifolia are probably conspecific. It seems best to treat the complex as a single polymorphic species. In historic times, human-assisted dispersal of possibly once distinct races has likely resulted in blurring of such distinctions with one exception. Plants of the outer coastal plain from Texas to North Carolina usually have a cluster of coarse hairs near the tip of phyllaries, at least in summer and fall-blooming plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 2. | FNA vol. 20, p. 234. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Heterotheca | Asteraceae > tribe Astereae > Heterotheca | ||||||||||||||||||||||||||||||||||||
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Synonyms | Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus | Inula subaxillaris | ||||||||||||||||||||||||||||||||||||
Name authority | (Pursh) Shinners: Field & Lab. 19: 71. (1951) | (Lamarck) Britton & Rusby: Trans. New York Acad. Sci. 7: 10. (1887) | ||||||||||||||||||||||||||||||||||||
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