Heterotheca villosa |
Heterotheca oregona |
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golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster |
Oregon false goldenaster, Oregon golden aster, rayless goldenaster |
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Habit | Perennials, (5–)16–40(–70) cm; taprooted. | Perennials, 15–105 cm; caudices branched. | ||||||||||||||||||||||||||||||||||||||||||||
Stems | 1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular. |
ascending to erect (sometimes ± brittle, sometimes brown), glabrate to densely hispid or densely stipitate-andular (axillary leaf fascicles sometimes present). |
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Leaves | generally not crowded; proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular. |
basal and proximal cauline withering by flowering; sessile; mid and distal blades ovate to linear-lanceolate, 12–50 × 4–40 mm, rounded to very weakly clasping, margins flat, strigoso-ciliate, apices acute, faces glabrate to densely hairy and/or stipitate-glandular; branch leaves reduced. |
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Peduncles | (4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular; bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads. |
5–50 mm, sparsely to densely hispid, stipitate-glandular; bracts linear to narrowly lanceolate, 3–9 × 0.5–2 mm. |
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Involucres | narrowly cylindric to campanulate, (5–)6–9.5(–13) mm. |
turbino-cylindric, 7.5–14 mm. |
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Ray florets | (5–)10–20(–38); laminae (3.5–)6.5–11(–20) × 1–2(–3) mm. |
0. |
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Disc florets | (10–)20–50(–85); corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm). |
(14–)20–45(–60); corollas ± ampliate, (7.5–)8.5–11(–14) mm, throats and lobes sparsely short-pilose, lobes 0.4–0.9 mm (margins thickened). |
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Phyllaries | in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular. |
in 5–7 series, outer ovate to deltate (2–5 × 0.6–1.7 mm), inner lanceolate, unequal, abaxial faces sparsely to densely hispid or stipitate-glandular (in green zone). |
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Heads | 1–16(–42), in usually open, corymbiform, rarely paniculiform arrays. |
(discoid) 1–15 per branch, in corymbiform, openly paniculiform, or racemiform arrays. |
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Cypselae | monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas). |
monomorphic, obconic, compressed, 2–5 × 0.35–1.6 mm, ribs 5–10, faces sparsely strigose; pappi tan, outer of linear scales 0.1–0.5 mm (not obvious), inner of 30–40 bristles 4.5–6.4, longest weakly clavate. |
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2n | = 18, 36. |
= 18. |
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Heterotheca villosa |
Heterotheca oregona |
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Distribution |
AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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CA; OR; WA
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Discussion | Varieties 9 (9 in the flora). Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
varieties 4 (4 in the flora Varieties 4 (4 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 2. | FNA vol. 20, p. 236. | ||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Heterotheca | Asteraceae > tribe Astereae > Heterotheca | ||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||
Synonyms | Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus | Ammodia oregona, Chrysopsis oregona | ||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Pursh) Shinners: Field & Lab. 19: 71. (1951) | (Nuttall) Shinners: Field & Lab. 19: 71. (1951) | ||||||||||||||||||||||||||||||||||||||||||||
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