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golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster

gray goldaster, hoary false goldenaster, hoary goldenaster

Habit Perennials, (5–)16–40(–70) cm; taprooted. Perennials, 15–40(–65) cm; taprooted, frequently rhizomatous (clonal).
Stems

1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular.

1–75, ascending to erect (proximaly sometimes reddish brown, often whitish due to pubescence, sometimes brittle), often short-branched in distal 1/2, usually sparsely long-hispid (more so in tetraploids), distally strigoso-canescent, eglandular (axillary leaf fascicles often present).

Leaves

generally not crowded;

proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular;

distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular.

generally ascending, congested;

proximal cauline petiolate or subsessile, blades oblanceolate, 19–32 × 2–6 mm, bases convex-cuneate to attenuate, margins flat, entire, strigoso-ciliate, proximally long-hispido-strigose, apices acute, faces densely strigose, long-hispid hairs few, eglandular;

distal sessile, usually silvery gray-green, blades linear-oblanceolate, 11–29 × 2–5 mm, little reduced distally, apices acute, faces very to extremely densely strigoso-canescent (90–200 hairs/mm²; silvery-whitish), long-hispid hairs usually few, eglandular.

Peduncles

(4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular;

bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads.

2–10 mm, densely strigose, usually sparsely hispid;

bracts grading from leaves, proximal oblanceolate, sometimes little reduced distally; long, linear-oblancelate, leaflike bracts often subtending heads.

Involucres

narrowly cylindric to campanulate, (5–)6–9.5(–13) mm.

cylindric to narrowly campanulate, 5–7(–7.5) mm (shorter than disc florets).

Ray florets

(5–)10–20(–38);

laminae (3.5–)6.5–11(–20) × 1–2(–3) mm.

10–22;

laminae 5–9(–10.5) × 0.8–1.4 mm.

Disc florets

(10–)20–50(–85);

corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm).

(14–)22–40(–50);

corollas ± ampliate, 4.7–6.5 mm, throats glabrous, lobes 0.5–0.75 mm, lobes sparsely pilose (hairs 0.1–0.25 mm).

Phyllaries

in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular.

in 4–6 series, linear-lanceolate to lanceolate, unequal (outer lengths 1/5–1/4 inner), margins scarious, distally ciliate and reddish purple, faces moderately strigose, eglandular.

Heads

1–16(–42), in usually open, corymbiform, rarely paniculiform arrays.

1–9(–15), borne singly or in corymbiform arrays, branches ascending.

Cypselae

monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose;

pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas).

monomorphic, obconic, compressed, 1.2–3.1 mm, ribs 6–10 (sometimes brownish), faces moderately strigose;

pappi off-white, outer of linear scales 0.25–0.5 mm, inner of 25–40 bristles 5–7.5 mm, longest weakly clavate.

2n

= 18, 36.

= 18, 36.

Heterotheca villosa

Heterotheca canescens

Phenology Flowering May–Jul(–Sep).
Habitat Sandy calcareous clay soils, igneous soils, sandy gypsiferous loamy soils, gravelly soils, outcrops, prairies, open hills, roadsides, fencerows
Elevation 100–1800 m [300–5900 ft]
Distribution
from FNA
AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
KS; NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 9 (9 in the flora).

Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Heterotheca canescens grows in southcentral Kansas, eastern New Mexico, western Oklahoma, and much of western Texas, with disjunct populations in the Davis and Glass mountains of trans-Pecos Texas. Disjunct collections putatively from Holt County, Missouri, are of questionable provenance. Heterotheca canescens is silvery gray-green due to the very dense indument of appressed hairs (usually more than 100 per mm2). Tetraploids have more hispid stems and more branched arrays than diploids and can be similar to forms of H. stenophylla var. angustifolia that grow sympatrically in west Texas and Oklahoma. The tetraploids of both taxa apparently hybidize, forming local swarms of parentlike and hybrid individuals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Asteraceae > tribe Astereae > Heterotheca Asteraceae > tribe Astereae > Heterotheca
Sibling taxa
H. camporum, H. canescens, H. fulcrata, H. grandiflora, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. stenophylla, H. subaxillaris, H. viscida, H. zionensis
H. camporum, H. fulcrata, H. grandiflora, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. stenophylla, H. subaxillaris, H. villosa, H. viscida, H. zionensis
Subordinate taxa
H. villosa var. ballardii, H. villosa var. depressa, H. villosa var. foliosa, H. villosa var. minor, H. villosa var. nana, H. villosa var. pedunculata, H. villosa var. scabra, H. villosa var. sierrablancensis, H. villosa var. villosa
Key
1. Stems sparsely to densely hispido-strigose, sparsely to densely stipitate-glandular; distal cauline leaves sparsely to densely hispido-strigose (hairs 2–100/mm²), sparsely to densely stipitate-glandular (glands 2–60/mm²)
→ 2
1. Stems moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular; distal leaf faces moderately to densely strigose (hairs 25–200/mm²), eglandular or sparsely stipitate-glandular (rarely more densely so)
→ 5
2. Distal cauline leaf blades ovate to oblanceolate or lanceolate to triangular-lanceolate, rarely oblong, glands sometimes more numerous than hairs/mm²; arrays corymbiform
→ 3
2. Distal cauline leaf blades linear, lanceolate, narrowly to broadly oblong, or narrowly ovate, glands less numerous than hairs/mm²; arrays congested, corymbiform to subumbelliform; w Great Plains, foothills of Front Range, White Mountains (New Mexico)
→ 4
3. Distal cauline leaf blades usually narrowly to broadly oblanceolate, sometimes oblong, sometimes ovate in mountains, margins flat to remotely undulate, apices obtuse to acute, indument sparse to moderate; arrays compact, branches usually not very long; phyllaries usually moderately strigose (hairs to 1 mm), often stipitate-glandular
var. minor
3. Distal cauline leaf blades narrowly lanceolate to triangular-lanceolate, apices sharply acute, margins flat; arrays lax, branches often very long; phyllaries sparsely short-strigose, hairs usually less than 0.3 mm, sparsely to moderately stipitate-glandular; rock outcrops, margins of arroyos, rocky banks, Arizona, Idaho, Nevada, Utah, disjunct in Little San Bernardino Mountains, California
var. scabra
4. Distal leaf blades often perpendicular to stems, linear to broadly oblong, sometimes small (less than 10 mm), faces moderately hispido-strigose (hairs 9–56/mm²), moderately to densely stipitate-glandular (glands 9–36/mm²), glands sometimes more than hairs; ray laminae (3.5–)6.9–11(–12) mm; lower elevations in mountains and foothills, Colorado and New Mexico, Great Plains, Colorado, w Nebraska, New Mexico, se Wyoming
var. nana
4. Distal leaf blades (perpendicular proximally, ascending distally) lanceolate to ovate-oblong, faces moderately to densely strigose (hairs usually 42–93/mm²), sparsely to moderately stipitate-glandular (glands usually 4–37/mm²); ray laminae (8.5–)9–12(–14) mm; mid elevations of Sierra Blanca, New Mexico, and mountains to s, w
var. sierrablancensis
5. Stems often abundantly long-hirsute; distal cauline leaf blades narrowly to broadly oblong; bracts subtending heads often surpassing them
→ 6
5. Stems sparsely to moderately long-hirsute; distal cauline leaf blades narrowly to broadly oblanceolate; bracts subtending heads small or none
→ 7
6. Leaf faces moderately strigose (hairs usually not obscuring faces); involucres broadly campanulate; rays showy, (12–)17–30 (averaging 21), laminae (8.5–)9.9–14.5 mm (averaging 11.5 mm); ne Great Plains
var. ballardii
6. Leaf faces moderately to densely hispido-strigose (hairs sometimes obscuring faces); involucres narrowly campanulate; rays less showy, (7–)13–22 (averaging 17), laminae 8–11.4(–12.4) mm (averaging 10 mm); mountains and w prairies
var. foliosa
7. Stems moderately to densely strigose, sparsely to abundantly long-hirsute; distal cauline leaf faces moderately strigose (20–60 hairs/mm²); Great Plains, scattered in hills and mountains
var. villosa
7. Stems sparsely long-hirsute; distal cauline leaf faces densely strigose (60–270 hairs/mm²)
→ 8
8. Stems 5–22 cm, sparsely long-hirsute; distal leaf faces densely strigose (84–140 hairs/mm²), sparsely to moderately stipitate-glandular; geyser basins, Yellowstone National Park, gravelly stream banks, e Teton National Park, rare further s in Wyoming
var. depressa
8. Stems (12–)30–50(–75) cm, usually sparsely, rarely abundantly long-hirsute; distal leaf faces densely strigose (60–270 hairs/mm²), eglandular or sparsely stipitate-glandular; grasslands, wooded hillsides, mountains, Arizona, Colorado, New Mexico, Utah
var. pedunculata
Synonyms Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus Haplopappus canescens, Chrysopsis berlandieri, Chrysopsis canescens, Chrysopsis villosa var. canescens
Name authority (Pursh) Shinners: Field & Lab. 19: 71. (1951) (de Candolle) Shinners: Field & Lab. 19: 68. (1951)
Source FNA vol. 20, p. 2. Treatment author: John C. Semple. FNA vol. 20, p. 246. Treatment author: John C. Semple.
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