Heterotheca villosa |
Heterotheca canescens |
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golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster |
gray goldaster, hoary false goldenaster, hoary goldenaster |
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| Habit | Perennials, (5–)16–40(–70) cm; taprooted. | Perennials, 15–40(–65) cm; taprooted, frequently rhizomatous (clonal). | ||||||||||||||||||||||||||||||||
| Stems | 1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular. |
1–75, ascending to erect (proximaly sometimes reddish brown, often whitish due to pubescence, sometimes brittle), often short-branched in distal 1/2, usually sparsely long-hispid (more so in tetraploids), distally strigoso-canescent, eglandular (axillary leaf fascicles often present). |
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| Leaves | generally not crowded; proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular. |
generally ascending, congested; proximal cauline petiolate or subsessile, blades oblanceolate, 19–32 × 2–6 mm, bases convex-cuneate to attenuate, margins flat, entire, strigoso-ciliate, proximally long-hispido-strigose, apices acute, faces densely strigose, long-hispid hairs few, eglandular; distal sessile, usually silvery gray-green, blades linear-oblanceolate, 11–29 × 2–5 mm, little reduced distally, apices acute, faces very to extremely densely strigoso-canescent (90–200 hairs/mm²; silvery-whitish), long-hispid hairs usually few, eglandular. |
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| Peduncles | (4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular; bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads. |
2–10 mm, densely strigose, usually sparsely hispid; bracts grading from leaves, proximal oblanceolate, sometimes little reduced distally; long, linear-oblancelate, leaflike bracts often subtending heads. |
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| Involucres | narrowly cylindric to campanulate, (5–)6–9.5(–13) mm. |
cylindric to narrowly campanulate, 5–7(–7.5) mm (shorter than disc florets). |
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| Ray florets | (5–)10–20(–38); laminae (3.5–)6.5–11(–20) × 1–2(–3) mm. |
10–22; laminae 5–9(–10.5) × 0.8–1.4 mm. |
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| Disc florets | (10–)20–50(–85); corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm). |
(14–)22–40(–50); corollas ± ampliate, 4.7–6.5 mm, throats glabrous, lobes 0.5–0.75 mm, lobes sparsely pilose (hairs 0.1–0.25 mm). |
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| Phyllaries | in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular. |
in 4–6 series, linear-lanceolate to lanceolate, unequal (outer lengths 1/5–1/4 inner), margins scarious, distally ciliate and reddish purple, faces moderately strigose, eglandular. |
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| Heads | 1–16(–42), in usually open, corymbiform, rarely paniculiform arrays. |
1–9(–15), borne singly or in corymbiform arrays, branches ascending. |
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| Cypselae | monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas). |
monomorphic, obconic, compressed, 1.2–3.1 mm, ribs 6–10 (sometimes brownish), faces moderately strigose; pappi off-white, outer of linear scales 0.25–0.5 mm, inner of 25–40 bristles 5–7.5 mm, longest weakly clavate. |
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| 2n | = 18, 36. |
= 18, 36. |
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Heterotheca villosa |
Heterotheca canescens |
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| Phenology | Flowering May–Jul(–Sep). | |||||||||||||||||||||||||||||||||
| Habitat | Sandy calcareous clay soils, igneous soils, sandy gypsiferous loamy soils, gravelly soils, outcrops, prairies, open hills, roadsides, fencerows | |||||||||||||||||||||||||||||||||
| Elevation | 100–1800 m [300–5900 ft] | |||||||||||||||||||||||||||||||||
| Distribution |
AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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KS; NM; OK; TX
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| Discussion | Varieties 9 (9 in the flora). Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Heterotheca canescens grows in southcentral Kansas, eastern New Mexico, western Oklahoma, and much of western Texas, with disjunct populations in the Davis and Glass mountains of trans-Pecos Texas. Disjunct collections putatively from Holt County, Missouri, are of questionable provenance. Heterotheca canescens is silvery gray-green due to the very dense indument of appressed hairs (usually more than 100 per mm2). Tetraploids have more hispid stems and more branched arrays than diploids and can be similar to forms of H. stenophylla var. angustifolia that grow sympatrically in west Texas and Oklahoma. The tetraploids of both taxa apparently hybidize, forming local swarms of parentlike and hybrid individuals. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Key |
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| Synonyms | Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus | Haplopappus canescens, Chrysopsis berlandieri, Chrysopsis canescens, Chrysopsis villosa var. canescens | ||||||||||||||||||||||||||||||||
| Name authority | (Pursh) Shinners: Field & Lab. 19: 71. (1951) | (de Candolle) Shinners: Field & Lab. 19: 68. (1951) | ||||||||||||||||||||||||||||||||
| Source | FNA vol. 20, p. 2. | FNA vol. 20, p. 246. | ||||||||||||||||||||||||||||||||
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